Kansas Herpetofaunal Atlas

An adult male from Russell County, Kansas. Image © Bob Ferguson.
An adult female from Russell County, Kansas. Image © Bob Ferguson.
Adult Eastern Collared Lizard from Kansas. Image © Suzanne L. Collins, CNAH.
Adult Collared Lizard from Russell County. © Aaerion Penguinatus.
REPTILIA (Reptiles) SQUAMATA (PART) (Other Lizards) CROTAPHYTIDAE (Collared and Leopard Lizards)

Eastern Collared Lizard
Crotaphytus collaris (Say in James, 1822: 252)
krō-tă-phī-tŭs — kŏl-lăr-ĭs

Conservation Status:
State: None
Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
 The Collared Lizard is characterized by four limbs, an ear opening on each side of the head, smooth granular scales on the back, and one or two dark bands on the neck. The body, large head, limbs, and tail are bluish green or green in males and light brown in females; both sexes have small dark and light spots. The neck has one or two black bands. The belly is white. The throat is yellow or orange in males. Males have larger heads, grow to a larger size, and are more brightly colored than females. During pregnancy, females (otherwise only shades of tan with brown blotches and white flecks dorsally) develop transverse orange bars along the sides of their body.
Adults normally 203- 306 mm (8-12 inches) in total length. The largest specimen from Kansas is a male (KU 84623) from Chase County with a snout-vent length of 110 mm and a total length of 302 mm (12 inches) collected by Charles J. Cole on 27 August 1963. The maximum length throughout the range is 355.6 mm (14 inches) (Conant and Collins, 1998).
Distribution:
 Common in association with rocky areas but may also substitute mammal burrows along canyons in the southwest and south-central portions of the state (first reported by Tihen and Sprague, 1939). In northcentral Kansas, it is found in association with outcrops of the Cretaceous Greenhorn Limestone, Dakota Formation, and Niobrara Chalk (rarely). In south-central Kansas is known from outcrops of Quaternary Ogallala, Cretaceous Kiowa Shale and Cheyenne Sandstone, and the various Permian groups. In southeastern Kansas (east of the Flint Hills, and south of the Kansas River) Eastern Collared Lizards have been found in association with outcrops of the Pennsylvanian Wabaunsee Group, Shawnee Group, Douglas Group, Lansing Group, Pleasanton Group, and the western edge of the Cherokee Group.
The specimens from Johnson (KU 189283), Kearney (221495), McPherson (KU 218857), Norton (FHSM 16932), and Stanton (KU 192220) counties are in need of corroboration. Subsequent visits to these sites have failed to yield additional specimens. They may represent individual releases.
Specimens exist for Hamilton County (KU 27746-8) but are only given to the county and therefore not plotted. It lends support to the Kearney and Stanton county records, as do specimens from adjacent populations in Colorado (Hammerson 1986,1999).
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
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Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 3,370
    Records 
  • 3,088
    Museum Vouchers 
  • 282
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
 County Breakdown: County Name (# occurrences):
 Allen (17); Anderson (141); Barber (113); Barton (4); Bourbon (73); Butler (22); Chase (315); Chautauqua (70); Cherokee (2); Clark (36); Clay (4); Cloud (2); Coffey (19); Comanche (25); Cowley (166); Crawford (75); Dickinson (20); Douglas (4); Elk (146); Ellis (212); Ellsworth (76); Ford (2); Geary (46); Greenwood (110); Hamilton (3); Harper (31); Johnson (2); Kearney (1); Kiowa (36); Labette (22); Lincoln (28); Linn (6); Logan (1); Lyon (25); Marion (33); Marshall (5); McPherson (1); Meade (84); Mitchell (18); Montgomery (62); Morris (6); Neosho (13); Ness (1); Norton (1); Osborne (1); Ottawa (9); Pottawatomie (60); Rice (1); Riley (356); Rooks (7); Rush (23); Russell (461); Saline (4); Sedgwick (1); Seward (6); Shawnee (3); Stanton (1); Sumner (1); Trego (5); Unknown (58); Wabaunsee (236); Washington (3); Wilson (52); Woodson (3);

Fossil History:
 Pleistocene fossil specimens are known from Meade County.
Fossils from the Cragin Quarry Local Fauna of Meade County (Pleistocene: Sangamonian) (Hay 1917; Etheridge 1958; Etheridge 1960, Tihen 1960; Tihen 1962; Brattstrom 1967; Preston 1979; Holman 1995) are assignable to this taxon.
Natural History:
 Collared Lizards are restricted to rocky areas on plains and near woodlands. Adults are active from April to September, although young individuals frequently are observed as late as mid-October. They prefer air temperatures ranging from 73° to 93°F. This aggressive, fast lizard is active only during the day, perching on rocks to bask in the sun and watch for enemies. If disturbed, it may run with its front feet held off the ground, using its strong hind limbs to race to a burrow or crevice beneath a large rock. Each Collared Lizard is territorial and has a home range of up to several thousand square feet. Invasion of a male's territory causes immediate response. The defending lizard displays his bright colors to discourage the invader. If this fails, the invader is attacked and chased from the area.
During winter months, Legler and Fitch (1957) found this species burrowed in small chambers beneath large rocks with small, partially dirt-filled tunnels leading to the edge of each rock. In these burrows the lizards were 203.2-304.8 mm (8-12 inches) below ground.
This lizard breeds during May and June after emerging from winter inactivity. Courtship involves the male displaying his bright throat and body colors to the female as he struts about her. After displaying, he moves around and over her in a relaxed, slithering walk, rapidly bobbing his head up and down. If the female is receptive, the male grasps the skin of her neck with his mouth, bends the rear part of his body under her tail, and copulation occurs.
Shortly after becoming pregnant and developing the characteristic orange-red spots or bars on her sides, the female seeks a nest of moist earth beneath a large rock or in a burrow. The eggs, numbering 2- 11 (Fitch, 1985), are laid within three weeks after mating and hatching occurs in two to three months. Females frequently have a second clutch each season.
Collared Lizards hunt their food by sight, running swiftly from their perches to grab observed prey. Grasshoppers, moths, beetles, spiders, wasps, and cicadas are eaten, as well as smaller lizards. Hallowell (1857) reported grasshoppers and beetles in the stomach of a specimen. Burt (1928) found that 95 percent of the food of Kansas specimens consisted of grasshoppers and moths.
Chief predators of the Collared Lizard are hawks and snakes. Werth (1974) reported aquatic behavior by this lizard in Ellis County.
Occurrence Activity:
 Number of Unique Obervations (=days): 200; Range: 02 Jan to 15 Nov
Remarks:
 First reported in Kansas by Hallowell (1856) from "Kansas". The earliest existing specimen (Museum of Comparative Zoology [MCZ 5937]) was collected at Fort Riley, Riley County. The MCZ catalog entry provides no other information, other than that the specimen was received from the Smithsonian Institution in 1879.
Eastern Collared Lizards can be difficult to capture. During cool weather, they can regularly be found snuggled up under rocks in a relatively torpid state where they are slow to move, and thus easy to pick up. However, during warm weather or sunny days, you’ll often need help; one person to lift a rock and the other to see which nearby rock it darted under. This process is repeated over and over (and typically involves lifting the same rocks several times) until one of you makes a successful lunge and pins the lizard against the ground. At this point, the collectors are either thankful for the leather gloves they’re wearing, or wishing that they would have thought ahead and put them on back at the car. The Eastern Collared Lizard is non-venomous and its teeth are actually quite small and of little consequence in its bite. The discomfort comes from the force of the bite; evolved to crush grasshoppers but effective at pinching unprotected fingers as well. Once controlled in hand, they can easily and safely be carried by grasping them around their body just behind the head. The lizard can then be placed temporarily in a pillowcase or other cloth bag for further examination or release.
Burt (1935) reported on a young collared lizard that was found under a rock in a prairie ledge near Altoona, Kansas, and in trying to escape it jumped into a large spider web from which it was unable to free itself without help.
Charles E. Burt (Burt and Hoyle 1935) chased a large Collared Lizard down a steep hill in Elk County, Kansas, on 22 April 1933, it dived into a pool of muddy water, where it hid in complete submergence beneath a small flat rock in an attempt to escape detection and capture
Bibliography:
1822 James, Edwin. Account of an expedition from Pittsburgh to the Rocky Mountains, performed in the years 1819 and ‘20, by order of the Hon. J. C. Calhoun, Sec’y of War: under the command of Major Stephen H. Long. From the notes of Major Long, Mr. T. Say, and other gentlemen of the exploring party. Compiled by Edwin James, botanist and geologist for the expedition. In two vols. – with an atlas. [Volume 2]. Henry Charles Carey and Isaac Lea, Philadelphia, Pennsylvania. 442, with supplementary chapters and an appendix.pp.
This is volume 2 (of 2) of the "Philadelphia" edition, which are dated 1823. However on 21 December 1822, Long sent a bound copy to John C. Calhoun (Secretary of War). It was publicly available, and effectively published for the sake of nomenclatural acts, on 31 December 1822. Woodman, 2010. Archives of Natural History 37(1): 28-38.
Contains the original description of Coluber testaceus (Masticophis flagellum testaceus) page 48, Ameiva tesselata (=Aspidoscelis tesselatus) page 50, Bufo cognatus (=Anaxyrus cognatus), Agama collaris (=Crotaphytus collaris) page 252, and Scincus lateralis (=Scincella lateralis) page 324, all by Thomas Say.
1856 Hallowell, Edward. Notice of a collection of reptiles from Kansas and Nebraska presented to the Academy of Natural Sciences, by Doctor Hammond, U. S. A. Proceedings of the Academy of Natural Sciences, Philadelphia 8():238-253
Contains reference to twenty-four species collected from 'Kansas', and includes the original description of Microps lineatus (=Tropidoclonion lineatum) from Kansas on page 241.
1859 Hallowell, Edward. Reports of Explorations and Surveys, to Ascertain the Most Practicable and Economical Route for a Railroad from the Mississippi River to the Pacific Ocean Vol. X, Part IV, No. 1, Washington, D. C. 1-27pp.
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():112-123
Also listed the Scarlet Snake (Cemophora coccinea) [=Ophibolus doliatus var. coccineus] and Red Salamander (Pseudotriton ruber) [=Spelerpes ruber] from Kansas.
1884 Garman, Samuel. The North American reptiles and batrachians. Bulletin of the Essex Institute 16():1-46
1897 Hurter, Julius. A contribution to the herpetology of Missouri. Transactions of the Academy of Science St. Louis 7(19):499-503
1900 Cope, Edward D. The crocodilians, lizards and snakes of North America. Pages 153-1270 in Report of the U. S. National Museum for the Year Ending June 30, 1898 , Washington, D. C. pp.
1911 Hurter, Julius. Herpetology of Missouri. Transactions of the Academy of Science St. Louis 20(5):59-274
1916 Householder, Victor H. The Lizards and Turtles of Kansas with Notes on Their Distribution and Habitat. Thesis. University of Kansas, Lawrence. 100pp.
1920 Taylor, Edward H. The Lizards of Kansas with Notes on Habits. Thesis. University of Kansas, Lawrence. 117pp.
Though Dr. Taylor's thesis lists 1916 as the publication date (as does version that the KHS published in 1993). His degree was not awarded until 1920, which is the official publication date.
1927 Burt, Charles E. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12
Accounts on forty species known from the vicinity of Manhattan, Riley County, Kansas. There have been several scientific names changes since publication, which is understandable... however, some usages cannot be explained by subsequent taxonomic reappraisals(e.g.  Gastrophryne carolinensis for Gastrophryne olivacea). The writer lists Eumeces (=Plestiodon) obsoletus and E. guttulatus yet correctly reasoning that the latter is the young of the former. Within the account of Tantilla gracilis (a common form) the writer mentions that T. nigriceps was reported from Riley County by Branson (1904) but that the specimens at Kansas State were absent at the time of publication. The Prairie Rattlesnake (Crotalus viridis) was first reported from Riley County by Branson (1904) and is reported by the writer based on a specimen in the museum at Kansas State University by Professor E. A. Popenoe. The Prairie Rattlesnake is not currently native to Riley County, and closest reliable localities are 150 to the west.
1928 Burt, Charles E. A key to the species of lizards definitely reported from Kansas. Privately printed, Enterprise Press, Bristow, Nebraska. pp.
Essentially a separate from the writer's "Lizards of. Kansas" which was in-press in the Transactions of the Academy of Science of St. Louis (Burt 1928. 26(1):1-81). Includes a glossary and a note on the variation in Sceloporus undulatus thayerii (= Sceloporus consobrinus).
1928 Ortenburger, Arthur I. The whip snakes and racers: Genera Masticophis and Coluber. Memiors of the University of Michigan Museum (1):1-247
1928 Burt, Charles E. The synonomy [sic], variation, and distribution of the collared lizard, Crotaphytus collaris (Say). Occasional Papers of the Museum of Zoology, University of Michigan (196):1-19
1928 Burt, Charles E. Insect food of Kansas lizards with notes on feeding habits. Journal of the Kansas Entomological Society 1(3):50-68
1928 Burt, Charles E. The lizards of Kansas. Transactions of the Academy of Science St. Louis 26(1):1-81
1929 Burt, Charles E. and May Danheim Burt. A collection of amphibians and reptiles from the Mississippi valley, with field observations. American Museum Novitates (381):1-14
1929 Burt, Charles E. The sexual dimorphism of the Collared Lizard, Crotaphytus collaris. Papers of the Michigan Academy of Science, Arts and Letters 10():417-421
1933 Stejneger, Leonhard and Thomas Barbour. A Checklist of North American Amphibians and Reptiles. 3rd Edition. Harvard University Press, Cambridge, Massachusetts. pp.
Reference to Kansas is the listed range of several species.
1933 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1935 Burt, Charles E. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16(3):311-366
1936 Hurd, Myron Alec. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103pp.
Under the supervision of thesis adviser Harry H. Hall. Report on 38 species (8 turtles, 7 lizards, and 23 snakes)... most unsubstantiated. Interesting inclusion are Crotalus horridus, Crotalus viridis, Kinosternon subrubrum, Opheodrys vernalis, and Phrynosoma cornutum.
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1939 Tihen, Joseph A. and James M. Sprague. Amphibians, reptiles, and mammals of the Meade County State Park Transactions of the Kansas Academy of Science 42():499-512
1940 Taylor, Edward H. Palatal sesamoid bones and palatal teeth in Cnemidophorus, with notes on these teeth in other saurian genera. Proceedings of the Biological Society of Washington 53():119-124
1941 Blair, W. Frank and Albert P. Blair. Food habits of the collared lizard in northeastern Oklahoma. American Midland Naturalist 26(1):230-232
1942 Rogers, Cornelius R. Observations on the natural control of Kansas termites with special reference to a bacterial disease affecting Reticulitermes tibialis Banks. ():
Noted feeding on swarming termites.
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1947 Hall, Henry H. and Hobart M. Smith. Selected records of reptiles and amphibians from southeastern Kansas Transactions of the Kansas Academy of Science 49(4):447-454
Report on certain Kansas specimens housed in the collection at what is now Pittsburg State University. Included are several species of dubious status today, including Cryptobranchus alleganiensis from the Neosho and Spring rivers (the only specimens from those significant drainages ever documented), Ambystoma maculatum from just north of Pittsburg, Crawford County, Heterodon nasicus from Crawford County, Opheodrys vernalis from Crawford County, Sonora episcopa from Crawford County, Agkistrodon piscivorus from Cherokee County, Crotalus atrox from Crawford County, and Crotalus viridis from Crawford County. They report several significant range extensions including Kinosternon flavescens from Turkey Creek in southeast Cherokee County, Graptemys geographica from just north of Pittsburg, Crotaphytus collaris from near Columbus, Cherokee County, Sceloporus consobrinus from just north of Pittsburg, Phrynosoma cornutum from Cherokee and Crawford counties, Heterodon platirhinos from Cherokee and Crawford counties, Haldea striatula from Crawford County, Sistrurus tergeminus from Crawford County, and a 402 lb Macrochelys temminckii in Cherokee County from just east of Chetopa (Labette County). They allude to the potential for Anaxyrus fowleri to occur in southeast Kansas and for native populations of Crotalus atrox in south central Kansas (in part from the disclosure that John R. Breukelman [then of ESU] had obtained three specimens in Woods County Oklahoma, 3/4 of a mile south of the Kansas line). None of the specimens the paper was based on, exist today.
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1951 Wolfenbarger, Keith. A. Systematic and Biological Studies on North American Chiggers of the genus Eutrombicula (Acarina, Trombiculidae). Thesis. University of Kansas, Lawrence. 77pp.
1951 Fitch, Henry S. and Wilmer W. Tanner. Remarks concerning the systematics of the collared lizard, Crotaphytus collaris, with a description of a new subspecies. Transactions of the Kansas Academy of Science 54(4):548-559
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Fitch, Henry S. An ecological study of the Collared Lizard (Crotaphytus collaris) University of Kansas Publications Museum of Natural History 8(3):213-274
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1957 Diener, Richard A. A western hognose snake eats a collared lizard. Herpetologica 13(2):122
1958 Etheridge, Richard E. Pleistocene lizards of the Cragin Quarry fauna of Meade County, Kansas. Copeia 1958(2):94-101
1960 Hibbard, Claude W. and Dwight W. Taylor. Two Late Pleistocene Faunas from southwestern Kansas. Contributions from the Museum of Paleontology, The University of Michigan 16(1):1-223
1960 Etheridge, Richard E. Additional notes on the lizards of the Cragin Quarry fauna. Papers of the Michigan Academy of Science, Arts and Letters 45():113-117
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1962 Robison, W. G .and Wilmer W. Tanner. A comparative study of the species of the genus Crotaphytus Holbrook (Iguanidae) Brigham Young University Science Bulletin 2(1):1–31
1964 Pleshkewych, A. Effects of Stimulation of the Motor Centers in the Cerebral Hemispheres of Pseudemys scripta and Crotaphytus collaris. Thesis. Emporia State University, Emporia, Kansas. 25pp.
1965 Clarke, Robert F. Lizards in Kansas. Kansas School Naturalist 11(4):1-16
1965 Weiner, Nancy J. and Hobart M. Smith. Comparative osteology and classification of the Crotaphytiform lizards. The American Midland Naturalist 73(1):170-187
1970 Webb, Robert G. Reptiles of Oklahoma. Stovall Museum, University of Oklahoma Press, Norman. 370pp.
1970 Fitch, Henry S. Reproductive cycles in lizards and snakes. University of Kansas Museum of Natural History Miscellaneous Publication (52):1-247
1972 Werth, Robert J. Lizard ecology: evidence of competition. Transactions of the Kansas Academy of Science 75():283-300
1973 Knight, James L., Eugene D. Fleharty, and Jerry D. Johnson. Noteworthy records of distribution and habits of some Kansas herptiles. Transactions of the Kansas Academy of Science 75(3):273-275
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Perry, Janice. KHS members take trip to southwest Kansas. Kansas Herpetological Society Newsletter (3):2-3
Account of a search for Crotalus atrox and other species discovered in Comanche County.
1974 Smith, Nathan M. and Wilmer W. Tanner. A taxonomic study of the western collared lizards, Crotaphytus collaris and Crotaphytus insularis. Brigham Young University Science Bulletin 19(4):1–29
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1975 Perry, Janice. A trip to southeastern Kansas. Kansas Herpetological Society Newsletter (7):4
1975 Capron, Marty B. A trip through the Kansas Flint Hills. Kansas Herpetological Society Newsletter (8):4-5
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Ferguson, Gary W. Color change and reproductive cycling in female Collared Lizards (Crotaphytus collaris). Copeia 1976():491-494
1977 Perry, Janice. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
Notable mentions: Spotted Salamander, Smooth Green Snake
1978 Skie, Shelley and Martha Bickford. KHS takes to the field in July at Winfield. Kansas Herpetological Society Newsletter (26):42798
1978 Capron, Marty B. Four county collecting raid: A south central Kansas herping saga. Kansas Herpetological Society Newsletter (26):9-12
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1979 Gray, Peter and Eddie Stegall. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8
1979 Guarisco, Hank. Historical note. Kansas Herpetological Society Newsletter (32):6-7
Report on the ephemeral population of Crotaphytus collaris in Douglas County, Kansas. Including predation records by Broad-winged Hawks.
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilis, Thamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1980 Spencer, Dwight. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64pp.
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1983 Fraser, John C. A trip to the 'TransPecos'. Kansas Herpetological Society Newsletter (54):18-23
1983 Miller, Larry L. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Heinrich, Mark L. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57pp.
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1984 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26
Invalidated the specimens of Thamnophis sirtalis from Hamilton County (reidentified as T. cyrtopsis; KU 2088) and Wallace County mapped in Collins, 1982.
1985 Sprackland, Robert G. Thoughts of a midwestern naturalist. Kansas Herpetological Society Newsletter (59):14-15
1985 Miller, Larry L. KHS 1985 field trip to Kirwin Reservoir. Kansas Herpetological Society Newsletter (61):11-12
1986 Capron, Marty B. In praise of the tiger. Kansas Herpetological Society Newsletter (63):12-13
1986 Olson, R. Earl, Bertram Marx, and Robert Rome. Descriptive dentition morphology of lizards of middle and North America, I: Scincidae, Teiidae, and Helodermatidae. Bulletin of the Maryland Herpetological Society 22(3):97-123
1987 Simmons, John E. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894
1987 Olson, R. Earl, Bertram Marx, and Robert Rome. Descriptive dentition morphology of lizards of middle and north America II: Iguanidae Bulletin of the Maryland Herpetological Society 23(1):12-34
1988 Bobbitt, Kitty Sue. Steroid Levels and Gravid Coloration in Crotaphytus collaris. Thesis. Emporia State University, Emporia, Kansas. 69pp.
1988 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19
1988 Miller, Larry L. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1988. Kansas Herpetological Society Newsletter (75):15-18
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Fitch, Henry S. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas. pp.
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1992 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Rundquist, Eric M. Results of the KHS 1992 fall field trip. Kansas Herpetological Society Newsletter (90):4
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1993 Sprackland, R. G. Husbandry and breeding of collared lizards. Vivarium 4(4):23-26
1993 Viets, Brian E. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113
1994 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1994 Riedle, J. Daren. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13
1995 Holman, J. Alan. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243pp.
1995 Anderson, Lewis, Mark Shaw, Jeff Blodig, and Tom Walker. Report to the Kansas Department of Wildlife and Parks: Herps encountered during REmap project, summer 1994. Kansas Herpetological Society Newsletter (99):10-17
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1996 Miller, Larry L. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Miller, Larry L. Many amphibian and reptile species identified during KHS 1996 fall field trip to Wabaunsee County. Kansas Herpetological Society Newsletter (106):2-3
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1996 McGuire, Jimmy A. Phylogenetic systematics of crotaphytid lizards (Reptilia: Iguania: Crotaphytidae). Bulletin of the Carnegie Museum of Natural History 32():iv + 1-143
1997 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1996. Kansas Herpetological Society Newsletter (107):14-16
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1997 Rundquist, Eric M. Addendum to 1997 KHS herp counts. Kansas Herpetological Society Newsletter (109):14-15
1997 Wells, W. Collared lizards of the genus Crotaphytus. (5(4)):48-68
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1998 Gamble, Jerre. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1998 Rundquist, Eric M. KDWP herp sting so far nets nine on Kansas and Federal charges. Kansas Herpetological Society Newsletter (112):5-6
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
1999 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1998. Kansas Herpetological Society Newsletter (116):14-15
2000 Taggart, Travis W. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5
2000 Taggart, Travis W. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Fogell, Daniel D. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6pp.
2002 Taggart, Travis W. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13
To Washington County, Kansas (also includes ancillary counts from Clay and Marshall counties).
2002 Mielke, E. Geographic distribution: Crotaphytus collaris. Journal of Kansas Herpetology (4):14
2002 Rundquist, Eric M. Natural history of the Night Snake, Hypsiglena torquata, in Kansas. Journal of Kansas Herpetology (4):16-20
2003 Fogell, Daniel D. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59pp.
This is the version the author submitted to the NPS. Their final publication was modified.
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 2002. Journal of Kansas Herpetology (5):13-16
2003 Platt, Dwight R. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (7):10
2003 Volkmann, Al. Cowley County herp count 1. Journal of Kansas Herpetology (7):7
2003 Lokke, John L. and Jill Lokke. Cowley County herp count 2. Journal of Kansas Herpetology (7):8
2003 Suleiman, Gibran. Fort Riley herp count. Journal of Kansas Herpetology (7):9
2004 Volkmann, Al. Cowley County herp count. Journal of Kansas Herpetology (11):10
2004 Collins, Joseph T. Marais des Cygnes herp count. Journal of Kansas Herpetology (11):11
2005 Taggart, Travis W. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Taggart, Travis W. Tiny T. rex: Collared Lizards in Kansas. Kansas Wildlife and Parks (October):15-20
2006 Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62():378-388
2007 Blevins, Emilie. Influence of landscape context on patterns of occupancy, abundance, and gene flow among Collared Lizards in the Flint Hills of Kansas. Thesis. Kansas State University, Manhattan, Kansas. 90pp.
2007 McGuire, Jimmy A., Charles W. Linkem, Michelle S. Koo, Delbert W. Hutchison, A. Kristopher Lappin, David I. Orange, Julio Lemos-Espinal, Brett R. Riddle, and Jef R. Jaeger. Mitochondrial introgression and incomplete lineage sorting through space and time: Phylogenetics of Crotaphytid lizards. Evolution 61-(12):2879–2897
2008 Taggart, Travis W. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3
2009 Murrow, Daniel G. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2010 Taggart, Travis W. Report on the KHS Fall Field Trip to Norton County, Kansas. Journal of Kansas Herpetology (36):8-10
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4
2011 Houck, Mike. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2012 Meshaka, Walter E., Jr., Stanley E. Trauth, Katherine M. Talbott, and Curtis J. Schmidt. Reproductive characteristics and sexual maturation of the Eastern Collared Lizard, Crotaphytus collaris (Say, 1823), at the northern edge of its geographic range. Collinsorum 1(2/3):9-14
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2012 Summer Field Trip to Meade County State Park. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Summer Field Trip to Coldwater Lake, Comanche County. Collinsorum 2(3/4):5
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2015 Mike Houck. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2015 Taggart, Travis W. Fall Field Trip Held In Washington County. Collinsorum 4(3):4
2016 Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. Vertebrate animals on the Fitch Natural History Reservation (1948-2002) Kansas Biological Survey Report Number 188, Lawrence. 48pp.
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3
2016 Taggart, Travis W. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2017 Taggart, Travis W. and J. Daren Riedle. A Pocket Guide to Kansas Amphibians, Turtles and Lizards. Great Plains Nature Center, Wichita, Kansas. 69pp.
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102
2017 Mardis, Dexter R. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10
2017 Taggart, Travis W. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8
2017 Schmidt, Curtis J. Herp Count: Southeast Ellis County. Collinsorum 6(2-3):9
2017 Taggart, Travis W. Herp Count: Northeast Barton County. Collinsorum 6(2-3):9
2018 Houck, Mike. Herp Count: Fort Riley Military Installation Collinsorum 7(1):17
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2020 Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150
2021 Mardis, Dexter R. Herp Count: Greenwood County: KHS-2020-14. Collinsorum 9(3):13
2021 Schmidt, Curtis J. Herp Count: Russell County: KHS-2020-16. Collinsorum 9(3):14
2021 Schmidt, Curtis J. Herp Count: Russell County: KHS-2020-19. Collinsorum 9(3):14
2021 Taggart, Travis W. Herp Count: Russell County: KHS-2020-24. Collinsorum 9(3):14-15
2023 Russell, Elisabeth Habitat associations and fine-scale movements of the Red-spotted Toad (Anaxyrus punctatus) in Kansas and the efficacy of remote telemetry for monitoring small-scale movements. Thesis. Fort Hays State University, Hays, Kansas. 81pp.
2024 Peacock, Daniel T. and Eva A. Horne. Characterization of mite parasitism of Eastern Collared Lizards (Crotaphytus collaris) in the Kansas Flint Hills. The Southwestern Naturalist 67(4):291-297
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Travis W. Taggart © 1999-2025 — w/ Sternberg Museum of Natural History, Fort Hays State University