1975
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Eshelman, Ralph E. Geology and paleontology of the early Pleistocene (late Blancan) White Rock fauna from northcentral Kansas. University of Michigan Museum of Palenontology, Papers on Paleontology. (13):60
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1979
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Van Devender, Thomas and Ralph E. Eshelman. Referral of the fossil lizard Sceloporus holmani (Late Pliocene of North-Central Kansas) to the genus Phrynosoma. Herpetologica 35(4):380-382
Sceloporus holmani from the late Pliocene of north-central Kansas, USA (Eschelman, 1975), isreferred to the genus Phrynosoma. The fossil record of Phrynosoma is discussed.
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1987
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Montanucci, Richard R. A phylogenetic study of the horned lizards, genus Phrynosoma, based on skeletal and external morphology. Natural History Museum of Los Angeles County, Contributions in Science (390):1-36
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1993
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Bell, Christopher J. Fossil lizards from the Elsinore Fault Zone, Riverside County, California. PaleoBios 15(2):18-26
Brief mention on the lack of knowledge regarding the squamosal horn patterns of P. adinognathus and P. holmani.
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