An adult from Meade County. Image © Suzanne L. Collins, CNAH.
An adult Western Ribbonsnake from Comanche County. © Maci Loughrea.
Adult from Wilson County. Image by Travis W. Taggart
An adult from Ness County. Image © Andrew Hare.
An adult from Jackson County. Image © Nick Gomez.
REPTILIA (Reptiles) SQUAMATA (PART) (Snakes) NATRICIDAE (Harmless Live-bearing Snakes)

Western Ribbonsnake
Thamnophis proximus (Say in James, 1822: 187)
thăm-nō-fĭs — prŏx-ĭ-mŭs


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
HARMLESS. The Western Ribbonsnake is characterized by keeled scales, a single anal scale, a stripe on each side of the body situated on the third and fourth scale rows (counting from the belly up), upper lips without dark vertical bars, and a uniform olive-gray, dark green, or black skin color between the stripes. The stripe down the middle of the back is red, orange, or yellow and those on each side of the body are light yellow or cream. The belly is white, cream, or gray with some dark coloration along its edges. Females grow slightly larger than males.
Adults normally attain 51.0- 76.0 mm (20-30 inches) in total length. The largest specimen from Kansas is a female (FHSM 7243) from Edwards County with a total length of 101.3 cm (39½ inches) collected by James Gubanyi and Keith Coleman on 2 June 2002. The maximum length throughout the range is 126.7 cm (49­7⁄8 inches) (Boundy, 1995; Powell et al., 2016).

Distribution:
Widespread in the eastern third of the state south of the Kansas River and largely confined to riparian corridors west of the Flint Hills.
Border vegetation around permanent bodies of water (swamps, marshes, ponds, rivers, ditches) and along the base of nearby rock outcrops where some individuals hibernate. Climbs onto piles of plants and low bushes.
Burt (1935) reported they are most abundant in extensive marshy areas that are filled with aquatic vegetation, such as reeds and cat-tails. They are often found in the reeds and grasses near shallow, meandering prairie streams as well, especially in the vicinity of those with sandy and gravelly bottoms.
Mozley (1877) first reported this species from Wallace County based on a specimen collected by Snow. This specimen no longer exists. Wilmer W. Tanner collected a specimen two miles east of Wallace (BYU 20742; no date is given). These are the westernmost reported specimens in Kansas, and additional corroborating specimens are desirable.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
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Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 790
    Records 
  • 715
    Museum Vouchers 
  • 75
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (11); Anderson (5); Atchison (1); Barber (55); Barton (3); Bourbon (5); Butler (9); Chase (20); Chautauqua (6); Cherokee (62); Clark (9); Coffey (4); Comanche (15); Cowley (6); Crawford (32); Doniphan (4); Douglas (49); Edwards (27); Elk (23); Ellis (42); Ellsworth (3); Finney (1); Ford (1); Franklin (37); Gove (1); Graham (4); Greenwood (17); Hamilton (1); Harper (11); Jackson (2); Jewell (1); Johnson (1); Kearney (3); Kingman (30); Kiowa (5); Labette (21); Leavenworth (1); Linn (7); Logan (1); Lyon (5); McPherson (2); Meade (47); Miami (13); Mitchell (1); Montgomery (12); Morris (3); Neosho (15); Ness (1); Osage (4); Osborne (1); Ottawa (1); Pawnee (1); Pratt (5); Reno (6); Rice (1); Riley (2); Rooks (1); Rush (1); Russell (13); Saline (5); Scott (9); Sedgwick (26); Seward (3); Shawnee (2); Sheridan (17); Stafford (14); Sumner (4); Trego (7); Unknown (5); Wabaunsee (6); Wallace (1); Washington (1); Wilson (1); Woodson (16); Wyandotte (2);

Fossil History:
Pleistocene fossil specimens are known from Meade and Jewell counties.
Fossils from the Williams Local Fauna of Rice County (Pleistocene: Illinoian) (Preston 1979; Holman 1984; Holman 1995) are assignable to this taxon.
Fossils from the Kanopolis Local Fauna of Ellsworth County (Pleistocene: Rancholabrean I) (Preston 1979, Holman 1972, Holman 1984; Holman 1995) are assignable to this taxon or T. sirtalis.
Fossils from the Mount Scott Local Fauna of Meade County (Pleistocene: Illinoian) (Preston 1979, Holman 1987; Holman 1995) are assignable to this taxon.
Fossils from the Butler Spring Local Fauna of Meade County (Pleistocene: Illinoian) (Tihen 1962; Preston 1979; Holman 1995) are assignable to this taxon.
Fossils from the Cragin Quarry Local Fauna of Meade County (Pleistocene: Sangamonian) (Hay 1917; Etheridge 1958; Etheridge 1960, Tihen 1960; Tihen 1962; Brattstrom 1967; Preston 1979; Holman 1995) are assignable to this taxon (species indeterminate).
Fossils from the Courtland Canal local fauna of Jewell County (Eshelman and Hager, 1984) are referable to this taxon.

Natural History:
This slender, nervous snake frequents the edges of swamps, marshes, lakes, streams, and rivers. It is active from March to October and is both diurnal and nocturnal depending upon prevailing temperatures. This species spends the day foraging along water-edge vegetation or basking in the sun on matted reeds and grasses. When frightened, it glides swiftly across water to escape. Gloyd (1928) found this species active and feeding on frogs at night during April in Franklin County. During winter, it retreats to burrows beneath the ground to avoid cold temperatures.
Western Ribbonsnakes mate during April and May, but courtship is unknown. Each female gives birth to young anytime from late June to September. Number of young per litter ranges from four to 28 (Fitch, 1985), with an average of twelve or thirteen. Gloyd (1928) collected a female in Franklin County which gave birth to eight young on 27 July. In Miami County, Gloyd (1932) found a female containing six well-developed young on 31 August. Collins (1974) obtained a gravid female collected in Franklin County which died on 13 August and contained twelve well-developed young.
This species feeds on small frogs, toads, salamanders, and fishes. Caldwell and Glass (1976) observed a Western Ribbonsnake eating a small Bullfrog in Woodson County. Taggart (1992) observed a Western Ribbonsnake consuming a Western Narrow-mouthed Toad in Chase County.
Predators of adult Western Ribbonsnakes include birds, mammals, and larger snakes. Young snakes are eaten by large frogs and fish (Collins, 1993).

Occurrence Activity:
Number of Unique Obervations (=days): 194; Range: 28 Feb to 16 Nov
Remarks:
First reported in Kansas by Mozley (1877) based on a specimen collected by KU entomologist Francis A. Snow in Wallace County. The earliest existing specimen (KU 2068) was collected in Clark County in May of 1903.
In regions with extensive upland grasslands in the western two-thirds of the state, this taxon can be found well away from streams and ponds.
Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity for this snake of three years, seven months, and one day.

Bibliography:
1822 James, Edwin. Account of an expedition from Pittsburgh to the Rocky Mountains, performed in the years 1819 and ‘20, by order of the Hon. J. C. Calhoun, Sec’y of War: under the command of Major Stephen H. Long. From the notes of Major Long, Mr. T. Say, and other gentlemen of the exploring party. Compiled by Edwin James, botanist and geologist for the expedition. In two vols. – with an atlas. [Volume 1]. Henry Charles Carey and Isaac Lea, Philadelphia, Pennsylvania. 503pp.
This is volume 1 (of 2) of the "Philadelphia" edition, which are dated 1823. However on 21 December 1822, Long sent a bound copy to John C. Calhoun (Secretary of War). It was publicly available, and effectively published for the sake of nomenclatural acts, on 31 December 1822. Woodman, 2010. Archives of Natural History 37(1): 28-38.
Contains the original description of Coluber obsoletus (=Pantherophis obsoletus) page 140, Coluber flaviventris (=Coluber constrictor flaviventris) page 185,  Coluber parietalis (Thamnophis sirtalis parietalis) page 186, Coluber proximus (=Thamnophis proximus) page 187, and Crotalus tergeminus (=Sistrurus tergeminus) page 499,  all by Thomas Say.
1877 Mozley, Annie E. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6():34-35
First attempt to compile a complete list of snakes reported from Kansas. Most specimens were from Douglas and Wallace counties.
Contains obvious identification/locality errors Thamnophis marcianus (Douglas County) and Heterodon simus (Wallace County).
Neroida holbrookii = Nerodia rhombifer.
Use of the name 'Kansas State University' actually refers to The University of Kansas where Annie E. (Mozley)Boddington received her BS @ KU in 1878 (see also Gloyd 1928).
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():112-123
Also listed the Scarlet Snake (Cemophora coccinea) [=Ophibolus doliatus var. coccineus] and Red Salamander (Pseudotriton ruber) [=Spelerpes ruber] from Kansas.
1882 Yarrow, Henry C. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249
A summary of all herpetological species known at the time, with reference to specimens in the United States National Museum. Including one three Acris blanchardi from Fort Riley; Agkistrodon contortrix from Fort Riley; three Ambystoma mavortium from "Kansas" and another from Fort Riley; one Anaxyrus woodhousii from "Kansas"; one Anaxyrus cognatus from "Kansas" and another from Fort Riley; one Carphophis vermis from Fort Scott; three Coluber constrictor from "Kansas" and two from Fort Riley; one Crotalus horridus from 1858; one Diadophis punctatus from Hyatt [Hyette sic], Kansas (Anderson County); one Graptemys pseudogeographica from the Republican River in Kansas;   two Heterodon nasicus from Fort Riley; one Lampropeltis calligaster from Neosho Falls; one Lampropeltis holbrooki from Fort Riley, one from "Natchez", Kansas, and one other from Shawnee Mission, Kansas;one Lampropeltis gentilis from Fort Riley and one other from the Republican River, Kansas; one Pantherophis obsoletus from Fort Riley;fourteen Phrynosoma douglassi from "Kansas" and four from Fort Riley; three Phrynosoma cornutum from Fort Riley (Riley County);  three Pituophis catenifer from "Platte Valley", Kansas [likely from eastern Colorado prior to 1861] and two specimens from Fort Riley; one Plestiodon septentrionalis from Neosho Falls (Woodson County); one Plestiodon obsoletus from Fort Riley; one Thamnophis sirtalis from "Kansas"; one Nerodia sipedon from Fort Riley and another from Neosho Falls; one Scincella lateralis from Fort Scott (Bourbon County); one Thamnophis proximus from Fort Riley; four Sceloporus consobrinus from Fort Riley; one Tantilla nigriceps from Fort Riley; four Thamnophis sirtalis from "Kansas" and two from Little Blue River, Kansas; 
1884 Garman, Samuel. The North American reptiles and batrachians. Bulletin of the Essex Institute 16():1-46
1885 Cragin, Francis W. Recent additions to the list of Kansas reptiles and batrachians, with further notes on species previously reported. Bulletin of the Washburn College Laboratory of Natural History 1(3):100-103
1885 Cragin, Francis W. Second contribution to the herpetology of Kansas, with observations on the Kansas fauna. Transactions of the Kansas Academy of Science 9():136-140
1903 Branson, Edwin B. Snakes of Kansas Thesis. University of Kansas, Lawrence. 41pp.
Describes all snake species reported from Kansas, considering doubtful those species that the author has not encountered himself. This list contains twenty-nine valid species. Collections examined included State University (KU), State House, Washburn College (WU), Ottawa University, State Normal School (ESU), State Agricultural College (KSU), and several high schools in Kansas. Examined the material available to Mozley (1878) and determined that of the thirty-three species listed, only twenty-three species were valid currently. And that Cragin's (1880) list of thirty-two species included eight specimens on the authority of Mozley that this author could not verify in any collection.
1904 Branson, Edwin B. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430
1908 Ruthven, Alexander G. Variations and genetic relationships of the garter-snakes. Bulletin of the United States National Museum (61):1-201
1925 Linsdale, Jean M. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312pp.
1927 Linsdale, Jean M. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81
1929 Taylor, Edward H. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62
1932 Gloyd, Howard K. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Museum of Zoology, University of Michigan 15():389-408
First record of Notophthalmus viridescens from Kansas. Second record (after the type locality) of Pseudacris crucifer from Kansas.
1933 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1935 Burt, Charles E. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16(3):311-366
1936 Hurd, Myron Alec. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103pp.
Under the supervision of thesis adviser Harry H. Hall. Report on 38 species (8 turtles, 7 lizards, and 23 snakes)... most unsubstantiated. Interesting inclusion are Crotalus horridus, Crotalus viridis, Kinosternon subrubrum, Opheodrys vernalis, and Phrynosoma cornutum.
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1937 Grant, Chapman. Herpetological notes from Central Kansas. American Midland Naturalist 18(3):370-372
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1939 Tihen, Joseph A. and James M. Sprague. Amphibians, reptiles, and mammals of the Meade County State Park Transactions of the Kansas Academy of Science 42():499-512
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1947 Hall, Henry H. and Hobart M. Smith. Selected records of reptiles and amphibians from southeastern Kansas Transactions of the Kansas Academy of Science 49(4):447-454
Report on certain Kansas specimens housed in the collection at what is now Pittsburg State University. Included are several species of dubious status today, including Cryptobranchus alleganiensis from the Neosho and Spring rivers (the only specimens from those significant drainages ever documented), Ambystoma maculatum from just north of Pittsburg, Crawford County, Heterodon nasicus from Crawford County, Opheodrys vernalis from Crawford County, Sonora episcopa from Crawford County, Agkistrodon piscivorus from Cherokee County, Crotalus atrox from Crawford County, and Crotalus viridis from Crawford County. They report several significant range extensions including Kinosternon flavescens from Turkey Creek in southeast Cherokee County, Graptemys geographica from just north of Pittsburg, Crotaphytus collaris from near Columbus, Cherokee County, Sceloporus consobrinus from just north of Pittsburg, Phrynosoma cornutum from Cherokee and Crawford counties, Heterodon platirhinos from Cherokee and Crawford counties, Haldea striatula from Crawford County, Sistrurus tergeminus from Crawford County, and a 402 lb Macrochelys temminckii in Cherokee County from just east of Chetopa (Labette County). They allude to the potential for Anaxyrus fowleri to occur in southeast Kansas and for native populations of Crotalus atrox in south central Kansas (in part from the disclosure that John R. Breukelman [then of ESU] had obtained three specimens in Woods County Oklahoma, 3/4 of a mile south of the Kansas line). None of the specimens the paper was based on, exist today.
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1951 Wolfenbarger, Keith. A. Systematic and Biological Studies on North American Chiggers of the genus Eutrombicula (Acarina, Trombiculidae). Thesis. University of Kansas, Lawrence. 77pp.
1951 Brumwell, Malcolm J. An ecological survey of the Fort Leavenworth Military Reservation American Midland Naturalist 45(1):187-231
Published posthumously. Lieutenant Brumwell died December 14, 1941, as a result of injuries incurred during the Japanese attack on Pearl Harbor. This paper is a condensed version of his thesis for the Master's degree.
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1960 Steward, P. L. Lung-flukes of snakes, genera Thamnophis and Coluber, in Kansas. University of Kansas Science Bulletin 41():877-890
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1962 Rossman, Douglas A. Thamnophis proximus (Say), a valid species of garter snake. Copeia 1962(4):741-748
1963 Rossman, Douglas A. The colubrid snake genus Thamnophis: A revision of the sauritus group. Bulletin of the Florida State Museum, Biological Sciences 7(3):99-178
1967 Choate, Jerry R. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46pp.
1967 Brattstrom, Bayard H. A succession of Pliocene and Pleistocene snake fauna from the High Plains of the United States. Copeia 1967(1):188-202
An examination (or in many cases, a re-examination) of 11,000 accumulated skeletal elements from fossil deposits collected at various sites and ranging in age from Lower Pliocene to the Recent.
1970 Rossman, Douglas A. Thamnophis proximus. Catalogue of American Amphibians and Reptiles (98):1-3
1972 Holman, J. Alan. Herpetofauna of the Kanopolis local fauna (Pleistocene: Yarmouth) of Kansas. Michigan Academic 5():87-98
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1975 Rundquist, Eric M. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3pp.
Short accounts for twenty-nine recognized amphibians and reptiles from Kingman County, Kansas. With habitat descriptions and for some species, estimates of population density.
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1976 Grow, David. The KHS goes to Chetopa. Kansas Herpetological Society Newsletter (13):2-3
Spring field trip (22 May 1976) along the Neosho River.
1976 Miller, Larry L. KHS visits Elk County. Kansas Herpetological Society Newsletter (15):1-2
1977 Perry, Janice. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
Notable mentions: Spotted Salamander, Smooth Green Snake
1978 Perry, Janice. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1979 Gray, Peter and Eddie Stegall. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilisThamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1980 Spencer, Dwight. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64pp.
1980 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survery of Kansas 9():1-11
1981 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survery of Kansas 10():7-19
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1982 Rogers, Karel L. Herpetofaunas of the Courland Canal and Hall Ash Local Faunas (Pleistoncene: Early Kansas) of Jewell Co., Kansas. Journal of Herpetology 16(2):174-177
1983 Simmons, John E. Fan fare. Kansas Herpetological Society Newsletter (54):16-18
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1984 Holman, J. Alan. Herpetofaunas of the Duck Creek and Williams Local Faunas (Pleistocene: Illinoian) of Kansas. Pages 20-38 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. pp.
1984 Eshelman, Ralph and Michael Hager. Two Irvingtonian (Medial Pleistocene) vertebrate faunas from northcentral Kansas. Pages 384-404 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8 Special Publication Number 8, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. pp.
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Heinrich, Mark L. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57pp.
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1984 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26
Invalidated the specimens of Thamnophis sirtalis from Hamilton County (reidentified as T. cyrtopsis; KU 2088) and Wallace County mapped in Collins, 1982.
1984 Collins, Joseph T. New records of fishes, amphibians and reptiles in Kansas for 1984. Kansas Herpetological Society Newsletter (58):14-20
1985 Lynch, John D. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Science 13():33-57
1986 Terry, P. A. Biological survey of the KS segments of Spring River and Shoal Creek. Part 1. Field Survey. Draft. Kansas Fish and Game, Pratt, Kansas. 67pp.
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1986 Holman, J. Alan. Butler Spring herpetofauna of Kansas (Pleistocene: Illinoian) and its climatic significance. Journal of Herpetology 20(4):568-569
1987 Holman, J. Alan. Climatic significance of a late Illinoian herpetofauna from southwestern Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(5):129-141
Anura - Bufo sp., ?Acris sp., Rana pipiens complex
Testudines - Sternotherus odoratus (Latreille), Chelydra serpentina (Linnaeus), Chrysemys picta (Schneider), Emydoidea blandingii (Holbrook), *Pseudemys hibbardi (Preston), Pseudemys scripta (Schoepff), Terrapene carolina (Linnaeus), Trionyx sp.
Squamata - Ophisaurus attenuatus Baird, Heterodon sp., Diadophis punctatus (Linnaeus), Coluber cf. C. constrictor Linnaeus, Elaphe vulpina (Baird and Girard), Lampropeltis getulus (Linnaeus), Pituophis melanoleucus (Daudin), Nerodia sipedon (Linnaeus), Regina grahami Baird and Girard, Storeria cf. S. dekayi, Thamnophis proximus (Say), Thamnophis radix (Baird and Girard), and Crotalinae indet were recovered.
1987 Holman, J. Alan. Snakes from the Robert Local Fauna (Late Wisconsinan) of Meade County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(6):143-150
Heterodon sp. indet, Diadophis punctatus (Linnaeus) Coluber constrictor Linnaeus, Lampropeltis getulus (Linnaeus) Nerodia sipedon (Linnaeus) Regina grahamii, Baird and Girard, Storeria cf. Storeria dekayi (Holbrook), Thamnophis proximus (Say), Thamnophis proxirnus (Say) or Thamnophis sirtalis (Linnaeus), and Thamnophis radix (Baird and Girard) recovered.
1988 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19
1988 Miller, Larry L. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1992 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17
1992 Rundquist, Eric M. Results of the KHS 1992 fall field trip. Kansas Herpetological Society Newsletter (90):4
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Viets, Brian E. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113
1993 Fitch, Henry S. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224
1994 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1995 Holman, J. Alan. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243pp.
1995 Cink, Calvin L. Seasonal activity of snakes on a Kansas floodplain tallgrass prairie. Proceedings of the 14th North American Prairie Conference, Kansas State University, Manhattan ():83-86
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1996 Rossman, Douglas A., Neil B. Ford, and Ricahrd A. Seigel. The Garter Snakes: Evolution and Ecology. University of Oklahoma Press, Norman. pp.
1996 Miller, Larry L. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Miller, Larry L. Many amphibian and reptile species identified during KHS 1996 fall field trip to Wabaunsee County. Kansas Herpetological Society Newsletter (106):2-3
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1997 Collins, Joseph T. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1998 Gamble, Jerre. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
1999 Taggart, Travis W. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6
Reported Anaxyrus woodhousii was likely A. fowleri.
2000 Taggart, Travis W. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5
2000 Taggart, Travis W. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2000 Taggart, Travis W. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8
2001 Taggart, Travis W. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Fogell, Daniel D. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6pp.
2002 Riedle, J. Daren and A. Hynek. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20
2002 Ellis, Mark R. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5
2002 Gubanyi, James E. and Keith Coleman. Size maxima: Thamnophis proximus. Journal of Kansas Herpetology (4):14
2002 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (4):15
2003 Freeman, Craig C. A natural areas inventory of the Ft. Leavenworth Military Reservation, Leavenworth County, Kansas. II. Open-file Report No. 117. Kansas Biological Survey, Lawrence, Kansas. 199pp.
2003 Fogell, Daniel D. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59pp.
This is the version the author submitted to the NPS. Their final publication was modified.
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 2002. Journal of Kansas Herpetology (5):13-16
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Gubanyi, James E. Western Ribbon Snake reproduction. Journal of Kansas Herpetology (7):12
2003 Volkmann, Al. Cowley County herp count 1. Journal of Kansas Herpetology (7):7
2003 Taggart, Travis W. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15
2004 Delisle, Jennifer M. and William H. Busby. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Volkmann, Al. Cowley County herp count. Journal of Kansas Herpetology (11):10
2004 Murrow, Daniel G. Geographic distribution. Thamnophis proximus. Kansas. Journal of Kansas Herpetology (11):14
County record, Wyandotte County, Kansas.
2005 Taggart, Travis W. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62():378-388
2008 Gomez, Nicholas J. Geographic distribution. Thamnophis proximus (Western Ribbon Snake). Journal of Kansas Herpetology (26):6
2009 Murrow, Daniel G. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769
2010 Miller, Larry L. Investigation of the Checkered Garter Snake in Kansas with notes on other Amphibians, Reptiles, and Turtles encountered. Kansas Department of Wildlife and Parks, Pratt, Kansas. 31pp.
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2014 McMartin, D. Chris. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2014 Taggart, Travis W. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12
2014 Taggart, Travis W. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2015 Taggart, Travis W. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3
2016 Taggart, Travis W. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):102
2017 Mardis, Dexter R. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10
2017 Taggart, Travis W. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2020 Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Neosho County: KHS-2020-01 Collinsorum 9(3):11
2021 Mardis, Dexter R. Herp Count: Sumner County: KHS-2020-08. Collinsorum 9(3):12-13
2021 Taggart, Travis W. Herp Count: Russell County: KHS-2020-24. Collinsorum 9(3):14-15
2021 Taggart, Megan M. and Travis W. Taggart. Herp Count: Seward County: KHS-2020-27. Collinsorum 9(3):15
2021 Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. Herp Count: Stafford County: KHS-2020-34. Collinsorum 9(3):16
2021 Riedle, J. Daren. Herp Count: Barton County: KHS-2020-28. Collinsorum 9(3):16
2021 Riedle, J. Daren. Herp Count: Pratt County: KHS-2020-33. Collinsorum 9(3):16
2022 Hallas, Joshua M., Thomas L. Parchman, and Chris R. Feldman. Phylogenomic analyses resolve relationships among garter snakes (Thamnophis: Natricinae: Colubridae) and elucidate biogeographic history and morphological evolution. Molecular Phylogenetics and Evolution 167():13
Utilized a genome-wide high-throughput sequencing data and near-complete taxon sampling to estimate a robust phylogeny for Thamnophis.
2023 Nunez, Leroy, Levi N Gray, David Weisrock, and Frank T Burbrink. The phylogenomic and biogeographic history of the Gartersnakes, Watersnakes, and Allies (Natricidae: Thamnophiini). Molecular Phylogenetics and Evolution 186(5:107844):12
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Travis W. Taggart © 1999-2024 — w/ Sternberg Museum of Natural History, Fort Hays State University