An adult Graham's Crawfish Snake from Barton County. © Maci Loughrea.
An adult Graham's Crawfish Snake from Greenwood County. © Jenny Smith.
 An adult Graham's Crawfish Snake from Cowley County. Image by Paxon Hutto.
An adult Graham's Crawfish Snake from Stafford County. Image © Eric Wood. iNat #70900267
REPTILIA (Reptiles) SQUAMATA (PART) (Snakes) NATRICIDAE (Harmless Live-bearing Snakes)

Graham's Crawfish Snake
Regina grahamii Baird & Girard 1853
rĕ-gī-nă — grăm-ī


Conservation Status:

State: None

Federal: None
NatureServe State: S4 - Apparently Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
HARMLESS. Characterized by keeled scales, a divided anal scale, and a distinct color and pattern. The head, body, and tail are a uniform brown or gray with a wide yellow or cream stripe running along each side near the belly. The belly is cream or white with a series of small dark spots. Adult males have slightly longer tails than females. Females grow larger than males.
The adults normally grow 45.7- 71.0 cm (18-28 inches) in total length. The largest specimen from Kansas is a female (KU 188175) from Lyon County with a total length of 102.9 cm (40­5⁄8 inches) collected by Robert F. Clarke on 11 June 1960. The maximum length throughout the range is 119.4 cm (47 inches) (Powell et al., 2016).

Distribution:
This occurs species along the margins of mud-bottom and rocky-edged marshes, oxbow lakes, rivers, and streams in areas with an abundance of crawfish.
Few records exist for northeastern Kansas (one each from Brown and Doniphan counties) but they are substantiated by records from southeastern Nebraska and northwestern Missouri.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
Open icons are questionable records; Click on a marker to view details.
Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 401
    Records 
  • 367
    Museum Vouchers 
  • 34
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (7); Anderson (6); Atchison (1); Barber (1); Barton (37); Bourbon (6); Brown (1); Butler (8); Chase (3); Chautauqua (3); Cherokee (12); Coffey (2); Cowley (11); Crawford (4); Dickinson (1); Douglas (72); Elk (2); Ellis (1); Ellsworth (1); Franklin (17); Geary (2); Greenwood (8); Harvey (11); Jewell (1); Johnson (10); Kingman (2); Labette (4); Linn (4); Lyon (12); Marshall (1); McPherson (2); Meade (3); Miami (9); Montgomery (2); Neosho (8); Osage (12); Pratt (10); Reno (2); Rice (3); Riley (3); Russell (1); Saline (3); Sedgwick (10); Shawnee (26); Stafford (27); Sumner (3); Unknown (2); Wabaunsee (2); Wilson (2); Woodson (20);

Fossil History:
Pleistocene fossil specimens are known from Meade, Rice, and Ellsworth counties. The Meade County records is outside the currently defined range of this species.
Fossils from the Williams Local Fauna of Rice County (Pleistocene: Illinoian) (Preston 1979; Holman 1984; Holman 1995) are assignable to this taxon.
Fossils from the Kanopolis Local Fauna of Ellsworth County (Pleistocene: Rancholabrean I) (Preston 1979, Holman 1972, Holman 1984; Holman 1995) are assignable to this taxon.
Fossils from the Mount Scott Local Fauna of Meade County (Pleistocene: Illinoian) (Preston 1979, Holman 1987; Holman 1995) are assignable to this taxon.
Fossils from the Butler Spring Local Fauna of Meade County (Pleistocene: Illinoian) (Tihen 1962; Preston 1979; Holman 1995) are assignable to this taxon.

Natural History:
Resides near ponds and sluggish streams of prairies, wet meadows, and river valleys. It is active from April to October. Like many other snakes, this species is active during the day in spring and fall and becomes nocturnal during the hot summer months. When not foraging for food, it retires into crawfish burrows to rest. During winter, this snake retreats into these same burrows to avoid cold temperatures. From 1 May to 6 June, Irwin and Collins (1987) recorded 82 Graham's Crawfish Snakes active at Cheyenne Bottoms in Barton County.
Probably breeds during April and May. Courtship occurs in the water and consists of preliminary frenzied swimming by several males, probably trying to locate a female. Eventually, the males discover a female and all may entwine with her, forming a fairly compact mass. More than one male may copulate with a single female.
Females give birth in late July, August, or September. Number of young per litter ranges from four to 39 (Fitch, 19856), with an average of sixteen. Males reach sexual maturity after one season, but females do not breed until their third year.
This snake feeds almost exclusively on crawfish (Collins, 1993).
Predators of this species include large birds, mammals, and other snakes.

Occurrence Activity:
Number of Unique Obervations (=days): 119; Range: 02 Jan to 28 Nov
Remarks:
First reported by Mozley (1877) based on a specimen collected by F. H. Morgan in Douglas County and deposited in the KU collection that existed at that time. Burt (1933) referenced a specimen in the US National Museum for Cairo (Pratt County) collected on 29 June 1892. However, the earliest existing specimen (KU 1305) from Kansas was collected in May of 1899 in Douglas County. The given locality of Harper County specimen (KU 186009) is actually in Barber County.
Additional surveys along the eastern sections of the Smoky Hill, Saline, Soloman, and Republican rivers may yield specimens.

Bibliography:
1877 Mozley, Annie E. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6():34-35
First attempt to compile a complete list of snakes reported from Kansas. Most specimens were from Douglas and Wallace counties.
Contains obvious identification/locality errors Thamnophis marcianus (Douglas County) and Heterodon simus (Wallace County).
Neroida holbrookii = Nerodia rhombifer.
Use of the name 'Kansas State University' actually refers to The University of Kansas where Annie E. (Mozley)Boddington received her BS @ KU in 1878 (see also Gloyd 1928).
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():112-123
Also listed the Scarlet Snake (Cemophora coccinea) [=Ophibolus doliatus var. coccineus] and Red Salamander (Pseudotriton ruber) [=Spelerpes ruber] from Kansas.
1885 Cragin, Francis W. Recent additions to the list of Kansas reptiles and batrachians, with further notes on species previously reported. Bulletin of the Washburn College Laboratory of Natural History 1(3):100-103
1885 Cragin, Francis W. Second contribution to the herpetology of Kansas, with observations on the Kansas fauna. Transactions of the Kansas Academy of Science 9():136-140
1904 Branson, Edwin B. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430
1927 Burt, Charles E. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12
Accounts on forty species known from the vicinity of Manhattan, Riley County, Kansas. There have been several scientific names changes since publication, which is understandable... however, some usages cannot be explained by subsequent taxonomic reappraisals(e.g.  Gastrophryne carolinensis for Gastrophryne olivacea). The writer lists Eumeces (=Plestiodon) obsoletus and E. guttulatus yet correctly reasoning that the latter is the young of the former. Within the account of Tantilla gracilis (a common form) the writer mentions that T. nigriceps was reported from Riley County by Branson (1904) but that the specimens at Kansas State were absent at the time of publication. The Prairie Rattlesnake (Crotalus viridis) was first reported from Riley County by Branson (1904) and is reported by the writer based on a specimen in the museum at Kansas State University by Professor E. A. Popenoe. The Prairie Rattlesnake is not currently native to Riley County, and closest reliable localities are 150 to the west.
1929 Dolman, Katherine. Studies of Kansas Water Snakes. Thesis. University of Kansas, Lawrence. 69pp.
1929 Taylor, Edward H. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62
1932 Gloyd, Howard K. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Museum of Zoology, University of Michigan 15():389-408
First record of Notophthalmus viridescens from Kansas. Second record (after the type locality) of Pseudacris crucifer from Kansas.
1933 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1936 Hurd, Myron Alec. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103pp.
Under the supervision of thesis adviser Harry H. Hall. Report on 38 species (8 turtles, 7 lizards, and 23 snakes)... most unsubstantiated. Interesting inclusion are Crotalus horridus, Crotalus viridis, Kinosternon subrubrum, Opheodrys vernalis, and Phrynosoma cornutum.
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1937 Grant, Chapman. Herpetological notes from Central Kansas. American Midland Naturalist 18(3):370-372
1942 Hudson, G. E. The amphibians and reptiles of Nebraska. Nebraska Conservation Bulletin 24():1-146
1947 Hall, Henry H. and Hobart M. Smith. Selected records of reptiles and amphibians from southeastern Kansas Transactions of the Kansas Academy of Science 49(4):447-454
Report on certain Kansas specimens housed in the collection at what is now Pittsburg State University. Included are several species of dubious status today, including Cryptobranchus alleganiensis from the Neosho and Spring rivers (the only specimens from those significant drainages ever documented), Ambystoma maculatum from just north of Pittsburg, Crawford County, Heterodon nasicus from Crawford County, Opheodrys vernalis from Crawford County, Sonora episcopa from Crawford County, Agkistrodon piscivorus from Cherokee County, Crotalus atrox from Crawford County, and Crotalus viridis from Crawford County. They report several significant range extensions including Kinosternon flavescens from Turkey Creek in southeast Cherokee County, Graptemys geographica from just north of Pittsburg, Crotaphytus collaris from near Columbus, Cherokee County, Sceloporus consobrinus from just north of Pittsburg, Phrynosoma cornutum from Cherokee and Crawford counties, Heterodon platirhinos from Cherokee and Crawford counties, Haldea striatula from Crawford County, Sistrurus tergeminus from Crawford County, and a 402 lb Macrochelys temminckii in Cherokee County from just east of Chetopa (Labette County). They allude to the potential for Anaxyrus fowleri to occur in southeast Kansas and for native populations of Crotalus atrox in south central Kansas (in part from the disclosure that John R. Breukelman [then of ESU] had obtained three specimens in Woods County Oklahoma, 3/4 of a mile south of the Kansas line). None of the specimens the paper was based on, exist today.
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1953 Schmidt, Karl P. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280pp.
Schmidt's first edition of his standardized checklist to North American amphibians and reptiles. Includes several specific references to Kansas in the range descriptions.
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1960 Cliburn, J. W. The Phylogeny and Zoogeography of North American Natrix. Dissertation. University of Alabama, Tuscaloosa. 319pp.
1960 Nakmura, E. L., and H. M. Smith. A comparative study of selected characters in certain American species of watersnakes . Transactions of the Kansas Academy of Science 63(2):102-113
1967 Choate, Jerry R. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46pp.
1969 Hall, Russell J. Ecological observations on Graham's Water Snake (Regina grahami Baird and Girard). American Midland Naturalist 81():156-163
1972 Holman, J. Alan. Herpetofauna of the Kanopolis local fauna (Pleistocene: Yarmouth) of Kansas. Michigan Academic 5():87-98
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1976 Grow, David. The KHS goes to Chetopa. Kansas Herpetological Society Newsletter (13):2-3
Spring field trip (22 May 1976) along the Neosho River.
1976 Capron, Marty B. and Jan Perry. A July weekend in Great Bend. Kansas Herpetological Society Newsletter (14):1-2
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1978 Hibbard, Claude W.; Richard J. Zakrzewski, Ralph E. Eshelman, Gordon Edmund, Clayton D. Griggs, and Caroline Griggs. Mammals from the Kanopolis Local Fauna, Pleistocene (Yarmouth) of Ellsworth County, Kansas. Contributions from the Museum of Paleontology, The University of Michigan 25(2):11-44
1979 Martin, Larry D. Survey of fossil vertebrates from east-central Kansas: Kansas River bank stabilization study. U.S. Army Corps of Engineers, Kansas City District. 55pp.
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilisThamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1980 Spencer, Dwight. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64pp.
1981 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survery of Kansas 10():7-19
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1984 Holman, J. Alan. Herpetofaunas of the Duck Creek and Williams Local Faunas (Pleistocene: Illinoian) of Kansas. Pages 20-38 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. pp.
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1984 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26
Invalidated the specimens of Thamnophis sirtalis from Hamilton County (reidentified as T. cyrtopsis; KU 2088) and Wallace County mapped in Collins, 1982.
1985 Lynch, John D. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Science 13():33-57
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1986 Holman, J. Alan. Butler Spring herpetofauna of Kansas (Pleistocene: Illinoian) and its climatic significance. Journal of Herpetology 20(4):568-569
1986 Cleveland, E. D. County record for Graham's Crayfish Snake (Regina grahamii). Transactions of the Kansas Academy of Science 89():9
1987 Holman, J. Alan. Climatic significance of a late Illinoian herpetofauna from southwestern Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(5):129-141
Anura - Bufo sp., ?Acris sp., Rana pipiens complex
Testudines - Sternotherus odoratus (Latreille), Chelydra serpentina (Linnaeus), Chrysemys picta (Schneider), Emydoidea blandingii (Holbrook), *Pseudemys hibbardi (Preston), Pseudemys scripta (Schoepff), Terrapene carolina (Linnaeus), Trionyx sp.
Squamata - Ophisaurus attenuatus Baird, Heterodon sp., Diadophis punctatus (Linnaeus), Coluber cf. C. constrictor Linnaeus, Elaphe vulpina (Baird and Girard), Lampropeltis getulus (Linnaeus), Pituophis melanoleucus (Daudin), Nerodia sipedon (Linnaeus), Regina grahami Baird and Girard, Storeria cf. S. dekayi, Thamnophis proximus (Say), Thamnophis radix (Baird and Girard), and Crotalinae indet were recovered.
1987 Holman, J. Alan. Snakes from the Robert Local Fauna (Late Wisconsinan) of Meade County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(6):143-150
Heterodon sp. indet, Diadophis punctatus (Linnaeus) Coluber constrictor Linnaeus, Lampropeltis getulus (Linnaeus) Nerodia sipedon (Linnaeus) Regina grahamii, Baird and Girard, Storeria cf. Storeria dekayi (Holbrook), Thamnophis proximus (Say), Thamnophis proxirnus (Say) or Thamnophis sirtalis (Linnaeus), and Thamnophis radix (Baird and Girard) recovered.
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1988. Kansas Herpetological Society Newsletter (75):15-18
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Seigel, Richard A. Ecology of a specialized predator: Regina grahami in Missouri. Journal of Herpetology 26(1):32-37
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1993 Viets, Brian E. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113
1993 Fitch, Henry S. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1994 Riedle, J. Daren. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13
1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4
See, Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.
1995 Holman, J. Alan. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243pp.
1995 Cink, Calvin L. Seasonal activity of snakes on a Kansas floodplain tallgrass prairie. Proceedings of the 14th North American Prairie Conference, Kansas State University, Manhattan ():83-86
1995 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1994. Kansas Herpetological Society Newsletter (100):24-47
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1998 Gamble, Jerre. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
2000 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1999. Kansas Herpetological Society Newsletter (119):7-9
2000 Taggart, Travis W. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2001 Taggart, Travis W. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2001 Alfaro, Michael E. and Stevan J. Arnold. Molecular systematics and evolution of Regina and the Thamnophiine snakes. Molecular Phylogenetics and Evolution 21(3):408-423
The T. lineatum used in their analyses was from Russell County, Kansas.
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Fogell, Daniel D. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6pp.
2002 Riedle, J. Daren and A. Hynek. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20
2002 Ernst, Carl H., J. Whitfield Gibbons and Michael E. Dorcas. Regina. Catalogue of American Amphibians and Reptiles (756):1-4
2003 Platt, Dwight R. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Schmidt, Curtis J. Geographic distribution: Regina grahamii. Journal of Kansas Herpetology (6):8
2003 Burr, Andrew. Coffey County herp count 1. Journal of Kansas Herpetology (7):7
2004 Gibbons, J. Whitfield and Michael E. Dorcas. North American Water Snakes. University of Oklahoma Press, Norman. pp.
2004 Collins, Joseph T. New records of amphibians, turtles, and reptiles in Kansas for 2003. Journal of Kansas Herpetology (9):8-11
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2008 Taggart, Travis W. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3
2009 Murrow, Daniel G. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769
2009 Collins, Joseph T. and Suzanne L. Collins. Geographic distribution. Regina grahamii. Journal of Kansas Herpetology (30):11
County record, Chase County, Kansas.
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):102
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2019 Anton, Thomas G. Regina grahamii. Catalogue of American Amphibians and Reptiles (920):1-40
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2021 Riedle, J. Daren. Herp Count: Barton County: KHS-2020-28. Collinsorum 9(3):16
2021 Tyson, Kelsea, Lexis Mader, Thomas Zapletal, Jeremiah Cline, Alyssa Farney, Loegan Hill, Jainee Cowen, Camron Matteson, and David Penning. Measuring herpetofaunal biodiversity in southwest Missouri. Collinsorum 10(2):13-18
2022 Ward, Krista J., Christine S. Streid, Jake T. Wright, and Thomas M. Luhring. Geographic distribution: Regina grahamii. USA, Kansas. Herpetological Review 53(2):264
2023 Nunez, Leroy, Levi N Gray, David Weisrock, and Frank T Burbrink. The phylogenomic and biogeographic history of the Gartersnakes, Watersnakes, and Allies (Natricidae: Thamnophiini). Molecular Phylogenetics and Evolution 186(5:107844):12
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