An adult Gophersnake from Hamilton County, Kansas. Image © Travis W. Taggart.
Image © Suzanne L. Collins, CNAH.
An juvenile Gophersnake from Bourbon County, Kansas. © Travis W. Taggart.
An adult Gophernake from Comanche County. © Maci Loughrea.
REPTILIA (Reptiles) SQUAMATA (PART) (Snakes) COLUBRIDAE (Harmless Egg-laying Snakes)

Gophersnake
Pituophis catenifer (Blainville 1835)
pĭt-ū-ō-fĭs — kă-tĭn-ĕh-fŭr


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
HARMLESS. The Gophersnake (aka Bullsnake) is unique among Kansas snakes in having an enlarged (but not upturned) rostral scale, keeled scales, and a single anal plate. The largest snake in Kansas, the Gophersnake is characterized by keeled scales, a single anal scale, and a pattern of 33- 73 large brown or black blotches on the brownish yellow body. The tail has alternating yellow and black bands. The belly is yellowish with variable black mottling. Adult males have longer tails than females.
The largest snake in Kansas. Adults normally grow 94.8-183.0 cm (37-72 inches) in total length. The largest specimen from Kansas is a female (KU 189258) from Harper County with total length of 226.0 cm (88­5⁄8 inches) and a weight of about 3,630 grams (8 pounds) collected by M. Kane on 21 May 1981. The maximum length throughout the range is 266.7 cm (105 inches) (Devitt et al., 2007; Powell et al., 2016).

Distribution:
The Gophersnake may be expected statewide but is generally only widespread west of the Flint Hills.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
Open icons are questionable records; Click on a marker to view details.
Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 2,189
    Records 
  • 1,461
    Museum Vouchers 
  • 728
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (13); Anderson (8); Atchison (1); Barber (85); Barton (16); Bourbon (15); Brown (2); Butler (18); Chase (15); Chautauqua (3); Cherokee (3); Cheyenne (23); Clark (47); Clay (7); Cloud (10); Coffey (3); Comanche (21); Cowley (51); Crawford (8); Decatur (11); Dickinson (10); Doniphan (8); Douglas (52); Edwards (14); Elk (12); Ellis (128); Ellsworth (22); Finney (263); Ford (50); Franklin (36); Geary (22); Gove (32); Graham (4); Grant (16); Gray (20); Greeley (3); Greenwood (17); Hamilton (50); Harper (47); Harvey (3); Haskell (7); Hodgeman (15); Jackson (2); Jefferson (3); Jewell (5); Johnson (9); Kearney (28); Kearny (8); Kingman (29); Kiowa (30); Labette (3); Lane (7); Leavenworth (2); Lincoln (3); Linn (3); Logan (52); Lyon (11); Marion (5); Marshall (27); McPherson (3); Meade (55); Miami (7); Mitchell (2); Montgomery (4); Morris (6); Morton (51); Nemaha (1); Ness (7); Norton (6); Osage (6); Osborne (12); Ottawa (2); Pawnee (17); Phillips (10); Pottawatomie (9); Pratt (22); Rawlins (13); Reno (43); Republic (6); Rice (9); Riley (42); Rooks (22); Rush (10); Russell (65); Saline (10); Scott (12); Sedgwick (33); Seward (48); Shawnee (4); Sheridan (5); Sherman (7); Smith (3); Stafford (45); Stanton (17); Stevens (12); Sumner (8); Thomas (7); Trego (66); Unknown (12); Wabaunsee (7); Wallace (16); Washington (19); Wichita (4); Wilson (2); Woodson (2); Wyandotte (2);

Fossil History:
Pleistocene fossil specimens are known from Meade, Ellsworth, and Rice counties.
Fossils from the Williams Local Fauna of Rice County (Pleistocene: Illinoian) (Preston 1979; Holman 1984; Holman 1995) are assignable to this taxon.
Fossils from the Kanopolis Local Fauna of Ellsworth County (Pleistocene: Rancholabrean I) (Preston 1979, Holman 1972, Holman 1984; Holman 1995) are assignable to this taxon.
Fossils from the Mount Scott Local Fauna of Meade County (Pleistocene: Illinoian) (Preston 1979, Holman 1987; Holman 1995) are assignable to this taxon.
Fossils from the Butler Spring Local Fauna of Meade County (Pleistocene: Illinoian) (Tihen 1962; Preston 1979; Holman 1995) are assignable to this taxon.

Natural History:
The Gophersnake lives in open grasslands as well as open woodland and woodland edge. It is common in cultivated fields where there is an abundance of rodents, its preferred food. Active from April to November at air temperatures above 60°F, it is generally diurnal, although Burt and Burt (1929) indicated that it may be active at night also. During the day, this snake basks in the sun or forages for food. Studies have shown that it uses its snout like a spade to shovel loose soil into a curve of its head and neck, after which the soil is dumped posteriorly away from the excavation; this activity may be useful in digging retreats or searching for food. With the approach of winter, this serpent seeks out deep crevices on rocky hillsides (eastern Kansas) or the burrows of small mammals (western Kansas), where it normally remains inactive until spring. Parks (1969) recorded a Gophersnake active in Ellis County during February.
Collins (1974) observed a Gophersnake partially emerged from an underground burrow during a late November warm period in Douglas County. During an unseasonably mild spell of weather, Capron (1986) observed an example of this species active in late January near Oxford. It is possible that this snake opportunistically emerges from its winter retreat whenever temperatures are favorable.
Gophersnakes mate during April and May after emergence from winter inactivity. Courtship involves the male crawling along and over the female until he eventually rests almost entirely on top of her. During this process, the male exhibits jerking body movements. The female is passive, except for elevating and waving her tail. Just prior to copulation the male may seize the female with his mouth, biting her on the head or neck. He curls his tail beneath hers until their cloacal openings meet; copulation commences and may last over an hour.
Female Gophersnakes lay their eggs in soft earth beneath large rocks or logs. Number of eggs per clutch ranges from three to 22, with thirteen being the average in the Great Plains (Fitch, 1985).
Gloyd (1928) found twelve eggs in a nest on 30 August in Franklin County. Marr (1944) found a female in Meade County which contained eleven eggs on 3 June. Mehrtens (1952) recorded a female from Saline County which laid nine eggs on 27 June. Four of the eggs hatched on 22-25 August; the young were about 279.4 mm (11 inches) long. Collins (1982) recorded a captive female from Douglas County (collected by Stanley Roth) that laid nine eggs on 14 July. Five of these successfully hatched on 30 September. Eric M. Rundquist (pers. comm., 1989; Collins, 1993) recorded a clutch of ten eggs laid on 30 June by a female from Sedgwick County.
The Gophersnake is probably the most economically beneficial snake in Kansas, consuming large quantities of rodents and saving farmers from much grain loss. This snake eats pocket gophers, rats, mice, rabbits, and ground squirrels. It also occasionally eats birds and bird eggs. Scheffer (1911) details the predatory behaviors of the Gophersnake on the Plains Pocket Gopher (Geomys bursarius) in Kansas. It is a widely-held, but incorrect, belief that Gophersnakes consume venomous snakes. While a Gophersnake and a Prairie Rattlesnake may have a similar diet (small mammals) their only general interaction may be competition for prey items.
Predators of Gophersnakes include large carnivorous birds and mammals (Collins, 1993). Young Gophersnakes may also be eaten by larger snakes.

Occurrence Activity:
Number of Unique Obervations (=days): 224; Range: 29 Jan to 08 Dec
Remarks:
The Gophersnake was first reported in Kansas by Cope (1875) based on specimens in the United States National Museum. The earliest extant specimen (MCZ 5877, 177822) was collected at "Ft. Riley, Kan" and received by the MCZ, from the Smithsonian Institution, around 1879. The jar containing MCZ 5877 was found to also contain an untagged specimen on 4 March 1992. On 22 March 1993 the untagged specimen was cataloged as MCZ 177822 (as per the penciled in notes on the MCZ catalog).
Gophersnakes are commonly thought to prey on other snakes, especially rattlesnakes, however, there is no evidence to support this claim. Both Gophersnakes and Kansas' rattlesnakes prey on small mammals.
Parks (1969) reported a Gophersnake active on 21 February 1969 in Ellis County.
The Bullsnake (P. c. sayi) is a variety of Gophersnake (P. catenifer) found east of the Rocky Mountains and the Pecos River and west of the Mississippi River, from northern Mexico into Canada.
Emits a loud “hiss” when disturbed or frightened, by forcefully exhaling air across a cartilaginous ridge situated In front of the glottis.
Knight and Collins (1977) recorded forty-one Gophersnakes during a two-week period in Cheyenne County and considered it the most abundant snake in that county.
Thirty-three of their specimens were found killed by vehicles on highways or roads. Capron (1985) recorded 38 of these snakes killed by swathing (mowing) machines during agricultural harvests from May to mid-July in southern Kansas.
Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity for this snake of 33 years and ten months.

Bibliography:
1837 Schlegel, Hermann. Essai sur la Physionomie des Serpens. [Essay on the Physiognomy of Serpents.] M. H. Schonekat, Libraire-Editeur, Amsterdam, Netherlands.. 251pp.
1875 Cope, Edward D. Check-list of North American Batrachia and Reptilia; with a systematic list of the higher groups, and an essay on geographical distribution. Based on the specimens contained in the U.S. National Museum. Bulletin of the United States National Museum 1():1-104
The first reference to Anaxyrus fowleri (nomen nudum) (not in Kansas). List the following species from Kansas: Carphophiops vermis (= Carphophis vermis); Ophibolus calligaster (= Lampropeltis calligaster); Ophibolus doliatus and Ophibolus doliatus annulatus (= Lampropeltis gentilis/triangulum); Tropidonotus sipedon erythrogaster (= Nerodia erythrogaster); Opheosaurus ventralis (= Ophisaurus attenuatus); Coluber emoryi (= Pantherophis emoryi); Coluber vulpinus (= Pantherophis ramspotti); Pituophis sayi (= Pituophis catenifer); and Tropidoclonion lineatum
1877 Mozley, Annie E. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6():34-35
First attempt to compile a complete list of snakes reported from Kansas. Most specimens were from Douglas and Wallace counties.
Contains obvious identification/locality errors Thamnophis marcianus (Douglas County) and Heterodon simus (Wallace County).
Neroida holbrookii = Nerodia rhombifer.
Use of the name 'Kansas State University' actually refers to The University of Kansas where Annie E. (Mozley)Boddington received her BS @ KU in 1878 (see also Gloyd 1928).
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():112-123
Also listed the Scarlet Snake (Cemophora coccinea) [=Ophibolus doliatus var. coccineus] and Red Salamander (Pseudotriton ruber) [=Spelerpes ruber] from Kansas.
1882 Yarrow, Henry C. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249
A summary of all herpetological species known at the time, with reference to specimens in the United States National Museum. Including one three Acris blanchardi from Fort Riley; Agkistrodon contortrix from Fort Riley; three Ambystoma mavortium from "Kansas" and another from Fort Riley; one Anaxyrus woodhousii from "Kansas"; one Anaxyrus cognatus from "Kansas" and another from Fort Riley; one Carphophis vermis from Fort Scott; three Coluber constrictor from "Kansas" and two from Fort Riley; one Crotalus horridus from 1858; one Diadophis punctatus from Hyatt [Hyette sic], Kansas (Anderson County); one Graptemys pseudogeographica from the Republican River in Kansas;   two Heterodon nasicus from Fort Riley; one Lampropeltis calligaster from Neosho Falls; one Lampropeltis holbrooki from Fort Riley, one from "Natchez", Kansas, and one other from Shawnee Mission, Kansas;one Lampropeltis gentilis from Fort Riley and one other from the Republican River, Kansas; one Pantherophis obsoletus from Fort Riley;fourteen Phrynosoma douglassi from "Kansas" and four from Fort Riley; three Phrynosoma cornutum from Fort Riley (Riley County);  three Pituophis catenifer from "Platte Valley", Kansas [likely from eastern Colorado prior to 1861] and two specimens from Fort Riley; one Plestiodon septentrionalis from Neosho Falls (Woodson County); one Plestiodon obsoletus from Fort Riley; one Thamnophis sirtalis from "Kansas"; one Nerodia sipedon from Fort Riley and another from Neosho Falls; one Scincella lateralis from Fort Scott (Bourbon County); one Thamnophis proximus from Fort Riley; four Sceloporus consobrinus from Fort Riley; one Tantilla nigriceps from Fort Riley; four Thamnophis sirtalis from "Kansas" and two from Little Blue River, Kansas; 
1883 Davis, N. S. Jr. and Frank L. Rice. Descriptive catalogue of North American batrachia and reptilia, found east of Mississippi River. Illinois Natural History Survey Bulletin 1(5):71
1885 Cragin, Francis W. Recent additions to the list of Kansas reptiles and batrachians, with further notes on species previously reported. Bulletin of the Washburn College Laboratory of Natural History 1(3):100-103
1885 Cragin, Francis W. Second contribution to the herpetology of Kansas, with observations on the Kansas fauna. Transactions of the Kansas Academy of Science 9():136-140
1886 Ebbutt, Percy G. Emigrant Life in Kansas Swan Sonnenschein and Company, Paternoster Square, London. 237pp.
Life around their homestead, seven miles from Parkerville along the Neosho River, in Morris County, Kansas. A lot of tales so tall as to render doubt on the few plausible scenarios presented. References to the deafening sounds of toads, as well as the abundance of lizards (including chameleons), and snakes. Many references to hunting and killing snakes. Many references to 'rattlesnakes' however, we can't tell what species they are. Includes an illustration of a Texas Horned Lizard and four illustrations of individuals killing snakes.
1903 Branson, Edwin B. Snakes of Kansas Thesis. University of Kansas, Lawrence. 41pp.
Describes all snake species reported from Kansas, considering doubtful those species that the author has not encountered himself. This list contains twenty-nine valid species. Collections examined included State University (KU), State House, Washburn College (WU), Ottawa University, State Normal School (ESU), State Agricultural College (KSU), and several high schools in Kansas. Examined the material available to Mozley (1878) and determined that of the thirty-three species listed, only twenty-three species were valid currently. And that Cragin's (1880) list of thirty-two species included eight specimens on the authority of Mozley that this author could not verify in any collection.
1904 Branson, Edwin B. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430
1911 Scheffer, Theo H. Distribution, natural enemies and breeding habits of the Kansas Pocket Gopher. Transactions of the Kansas Academy of Science 23/24():109-114
1917 Wooster, Lyman D. Nature Study Bulletin Kansas State Printing Plant, Topeka, Kansas.. 63pp.
1925 Linsdale, Jean M. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312pp.
1927 Burt, Charles E. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12
Accounts on forty species known from the vicinity of Manhattan, Riley County, Kansas. There have been several scientific names changes since publication, which is understandable... however, some usages cannot be explained by subsequent taxonomic reappraisals(e.g.  Gastrophryne carolinensis for Gastrophryne olivacea). The writer lists Eumeces (=Plestiodon) obsoletus and E. guttulatus yet correctly reasoning that the latter is the young of the former. Within the account of Tantilla gracilis (a common form) the writer mentions that T. nigriceps was reported from Riley County by Branson (1904) but that the specimens at Kansas State were absent at the time of publication. The Prairie Rattlesnake (Crotalus viridis) was first reported from Riley County by Branson (1904) and is reported by the writer based on a specimen in the museum at Kansas State University by Professor E. A. Popenoe. The Prairie Rattlesnake is not currently native to Riley County, and closest reliable localities are 150 to the west.
1927 Pope, T. E. B. A two-headed bullsnake. Yearbook. Publication of the Museum of Milwaukee 1925(5):161-167
Report on a bicephalic Gophersnake collected in 1902 at Sylvan Grove (Lincoln County), Kansas.
1927 Linsdale, Jean M. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81
1929 Burt, Charles E. and May Danheim Burt. A collection of amphibians and reptiles from the Mississippi valley, with field observations. American Museum Novitates (381):1-14
1929 Taylor, Edward H. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62
1929 Taylor, Edward H. List of reptiles and batrachians of Morton County, Kansas, reporting species new to the state fauna. University of Kansas Science Bulletin 19(6):63-65
Annotated listing of 25 species discovered in southwest Morton County just prior to the Dust Bowl that wreaked havoc on the region and the subsequent creation of the Cimarron National Grassland. Of special note are Heterodon platirhinos, Thamnophis marcianus, and Anaxyrus debilis (all of which have not been found in the area since).
1932 Stull, Olive G. An annotated list of the forms of the genus Pituophis. Occasional Papers of the Museum of Zoology, University of Michigan (250):1-5
1932 Gloyd, Howard K. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Museum of Zoology, University of Michigan 15():389-408
First record of Notophthalmus viridescens from Kansas. Second record (after the type locality) of Pseudacris crucifer from Kansas.
1933 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1935 Burt, Charles E. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16(3):311-366
1936 Hurd, Myron Alec. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103pp.
Under the supervision of thesis adviser Harry H. Hall. Report on 38 species (8 turtles, 7 lizards, and 23 snakes)... most unsubstantiated. Interesting inclusion are Crotalus horridus, Crotalus viridis, Kinosternon subrubrum, Opheodrys vernalis, and Phrynosoma cornutum.
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1937 Grant, Chapman. Herpetological notes from Central Kansas. American Midland Naturalist 18(3):370-372
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1938 Bond, Glenn Carl Serological studies of the Reptilia. Dissertation. University of Kansas, Lawrence, Kansas. 110pp.
1938 Schmidt, Karl P. Herpetological evidence for the postglacial eastward extension of the steppe in North America. Ecology 19(3):396-407
1939 Tihen, Joseph A. and James M. Sprague. Amphibians, reptiles, and mammals of the Meade County State Park Transactions of the Kansas Academy of Science 42():499-512
1940 Stull, Olive G. Variations and relationships in the snakes of the genus Pituophis. Bulletin of the United States National Museum (175):1-225
1941 Branson, F. and C. Deyoe. A study of snakes and lizards of Ellis County. Unpublished data, Fort Hays State University. pp.
Have been unable to locate a copy. 
Cited in Gish (1961. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.) among the species accounts under "other specimens reported". TWT 11 February 2020.
1941 Schmidt, Karl Peterson and D. D. Davis. Field Book of Snakes of the United States and Canada. C.P. Putnam and Sons, New York. 365pp.
1942 Anderson, Paul. Amphibians and reptiles of Jackson County, Missouri. Bulletin of the Chicago Academy of Sciences 6(11):203-222
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1945 Bugbee, R. E. A note on the mortality of snakes on highways in western Kansas. Transactions of the Kansas Academy of Science 47():373-374
1947 Hall, Henry H. and Hobart M. Smith. Selected records of reptiles and amphibians from southeastern Kansas Transactions of the Kansas Academy of Science 49(4):447-454
Report on certain Kansas specimens housed in the collection at what is now Pittsburg State University. Included are several species of dubious status today, including Cryptobranchus alleganiensis from the Neosho and Spring rivers (the only specimens from those significant drainages ever documented), Ambystoma maculatum from just north of Pittsburg, Crawford County, Heterodon nasicus from Crawford County, Opheodrys vernalis from Crawford County, Sonora episcopa from Crawford County, Agkistrodon piscivorus from Cherokee County, Crotalus atrox from Crawford County, and Crotalus viridis from Crawford County. They report several significant range extensions including Kinosternon flavescens from Turkey Creek in southeast Cherokee County, Graptemys geographica from just north of Pittsburg, Crotaphytus collaris from near Columbus, Cherokee County, Sceloporus consobrinus from just north of Pittsburg, Phrynosoma cornutum from Cherokee and Crawford counties, Heterodon platirhinos from Cherokee and Crawford counties, Haldea striatula from Crawford County, Sistrurus tergeminus from Crawford County, and a 402 lb Macrochelys temminckii in Cherokee County from just east of Chetopa (Labette County). They allude to the potential for Anaxyrus fowleri to occur in southeast Kansas and for native populations of Crotalus atrox in south central Kansas (in part from the disclosure that John R. Breukelman [then of ESU] had obtained three specimens in Woods County Oklahoma, 3/4 of a mile south of the Kansas line). None of the specimens the paper was based on, exist today.
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1951 Wolfenbarger, Keith. A. Systematic and Biological Studies on North American Chiggers of the genus Eutrombicula (Acarina, Trombiculidae). Thesis. University of Kansas, Lawrence. 77pp.
1951 Brumwell, Malcolm J. An ecological survey of the Fort Leavenworth Military Reservation American Midland Naturalist 45(1):187-231
Published posthumously. Lieutenant Brumwell died December 14, 1941, as a result of injuries incurred during the Japanese attack on Pearl Harbor. This paper is a condensed version of his thesis for the Master's degree.
1952 Mehrtens, J. M. Notes on the eggs and young of Pituophis melanoleucus sayi. Herpetologica 8():69-70
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Diener, Richard A. New records of snakes in southwestern Kansas. The Southwestern Naturalist 1(1):27-29
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1958 Smith, Ronald E. Natural history of the Prairie Dog in Kansas. University of Kansas Museum of Natural History, Miscellaneous Publications (16):1-36
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1967 Choate, Jerry R. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46pp.
1967 Brattstrom, Bayard H. A succession of Pliocene and Pleistocene snake fauna from the High Plains of the United States. Copeia 1967(1):188-202
An examination (or in many cases, a re-examination) of 11,000 accumulated skeletal elements from fossil deposits collected at various sites and ranging in age from Lower Pliocene to the Recent.
1969 Parks, Leland H. An active Bull Snake in near-freezing temperature. Transactions of the Kansas Academy of Science 72(2):266
1972 Holman, J. Alan. Herpetofauna of the Kanopolis local fauna (Pleistocene: Yarmouth) of Kansas. Michigan Academic 5():87-98
1973 McLeran, V. Friendly constrictors. Kansas Fish and Game (30(2)):8-11
Popular article: states that "Kansas is host to six constrictor reptiles" (Lampropeltis calligaster, L. holbrookia, L. gentilis/triangulum, Pantherophis emoryi, Pantherophis obsoletus, and Pituophis catenifer)... omitting Arizona elegans and Rhinocheilus lecontei. The image of the 'milk snake' is actually an Eastern Kingsnake (Lampropeltis getula) and the bullsnake in the Red-tailed Hawk nest (apparently a natural predation event recorded by the author) is a Prairie Kingsnake (Lampropeltis calligaster).
1974 Eshelman, Ralph E. Geology and paleontology of the early Pleistocene Belleville Formation of north central Kansas. Dissertation. University of Michigan, Ann Arbor, Michigan. 137pp.
1974 Henderson, Robert W. Resource partitioning among snakes of the University of Kansas Natural History Reservation: A preliminary analysis. Milwaukee Public Museum Contributions in Biology and Geology (1):1-11
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Perry, Janice. KHS members take trip to southwest Kansas. Kansas Herpetological Society Newsletter (3):2-3
Account of a search for Crotalus atrox and other species discovered in Comanche County.
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1975 Rundquist, Eric M. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3pp.
Short accounts for twenty-nine recognized amphibians and reptiles from Kingman County, Kansas. With habitat descriptions and for some species, estimates of population density.
1975 Rundquist, Eric M. First KHS field trip yields three county records. Kansas Herpetological Society Newsletter (7):1-3
Narration of the activities and species found during the KHS field trip to Kingman County, Kansas. From the title of the article, there were three county records were obtained, however, only Plestiodon septentrionalis is indicated as being 'new'.
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1976 Capron, Marty B. and Jan Perry. A July weekend in Great Bend. Kansas Herpetological Society Newsletter (14):1-2
1976 Rickart, Eric A. A new horned lizard (Phrynosoma adinognathus) from the early Pleistocene of Meade County, Kansas, with comments on the herpetofauna of the Borchers locality. Herpetologica 32(1):64-67
Contains the original description of Phrynosoma adinognathus.
1977 Knight, James L. and Joseph T. Collins. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19pp.
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
Notable mentions: Spotted Salamander, Smooth Green Snake
1978 Warner, M. and R. Wencel. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16
1978 Schwaner, Terry D. KHS field trip to Grant County, Kansas, 12-14 May 1978. Kansas Herpetological Society Newsletter (25):3-4
1978 Capron, Marty B. Four county collecting raid: A south central Kansas herping saga. Kansas Herpetological Society Newsletter (26):9-12
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1978 Hibbard, Claude W.; Richard J. Zakrzewski, Ralph E. Eshelman, Gordon Edmund, Clayton D. Griggs, and Caroline Griggs. Mammals from the Kanopolis Local Fauna, Pleistocene (Yarmouth) of Ellsworth County, Kansas. Contributions from the Museum of Paleontology, The University of Michigan 25(2):11-44
1979 Martin, Larry D. Survey of fossil vertebrates from east-central Kansas: Kansas River bank stabilization study. U.S. Army Corps of Engineers, Kansas City District. 55pp.
1979 Ports, Mark A. Occurrence and density studies of nongame wildlife in southwestern Kansas - May 16-August 16, 1979. Kansas Fish and Game Commission, Pratt, Kansas. 83pp.
1979 Gray, Peter and Eddie Stegall. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8
1979 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1978. Technical Publication of the State Biological Survery of Kansas 8():56-66
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilisThamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1980 Spencer, Dwight. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64pp.
1980 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survery of Kansas 9():1-11
1981 Miller, Larry L. Record length Bullsnake from Harper County, Kansas. Kansas Herpetological Society Newsletter (43):4
1981 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survery of Kansas 10():7-19
1982 Fitch, Henry S. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C. pp.
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1983 Simmons, John E. Fan fare. Kansas Herpetological Society Newsletter (54):16-18
1983 Miller, Larry L. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1983 Halpin, Zuleyma T. Naturally occurring encounters between Blacktailed Prairie Dogs (Cynomys ludovicianus) and snakes. American Midland Naturalist 109(1):50-54
Details encounters occurring on a 1.7-ha prairie dog town located at the Quivira National Wildlife Refuge in south central Kansas.
1984 Holman, J. Alan. Herpetofaunas of the Duck Creek and Williams Local Faunas (Pleistocene: Illinoian) of Kansas. Pages 20-38 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. pp.
1984 Platt, Dwight R. Growth of Bullsnakes (Pituophis melanoleucus sayi) on a sand prairie in south central Kansas. Pages 41-55 in Vertebrate Ecology and Systematics— A Tribute to Henry S Fitch. Special Publication Number 10. The University of Kansas Museum of Natural History, Lawrence. pp.
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Heinrich, Mark L. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57pp.
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1984 Collins, Joseph T. New records of fishes, amphibians and reptiles in Kansas for 1984. Kansas Herpetological Society Newsletter (58):14-20
1985 Capron, Marty B. A western diamondback rattlesnake released in Sumner County, Kansas. Kansas Herpetological Society Newsletter (59):5-6
Report on a specimen of Crotalus atrox discovered near Belle Plain, Sumner County, Kansas.
1985 Miller, Larry L. KHS 1985 field trip to Kirwin Reservoir. Kansas Herpetological Society Newsletter (61):11-12
1985 Lynch, John D. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Science 13():33-57
1986 Capron, Marty B. Winter activity noted in southern Kansas herps. Kansas Herpetological Society Newsletter (64):15-16
1986 Brunson, Ken. Some unusual injuries to snakes. Kansas Herpetological Society Newsletter (65):13-14
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1986 Holman, J. Alan. Butler Spring herpetofauna of Kansas (Pleistocene: Illinoian) and its climatic significance. Journal of Herpetology 20(4):568-569
1987 Kizirian, David A. Geographic variation in the zone of  contact between two subspecies of Gopher Snakes (Pituophis catenifer). Thesis. University of Texas at El Paso, El Paso, Texas. 69pp.
1987 Miller, Larry L. Another Bullsnake story. Kansas Herpetological Society Newsletter (68):17-18
1987 Simmons, John E. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894
1987 Holman, J. Alan. Climatic significance of a late Illinoian herpetofauna from southwestern Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(5):129-141
Anura - Bufo sp., ?Acris sp., Rana pipiens complex
Testudines - Sternotherus odoratus (Latreille), Chelydra serpentina (Linnaeus), Chrysemys picta (Schneider), Emydoidea blandingii (Holbrook), *Pseudemys hibbardi (Preston), Pseudemys scripta (Schoepff), Terrapene carolina (Linnaeus), Trionyx sp.
Squamata - Ophisaurus attenuatus Baird, Heterodon sp., Diadophis punctatus (Linnaeus), Coluber cf. C. constrictor Linnaeus, Elaphe vulpina (Baird and Girard), Lampropeltis getulus (Linnaeus), Pituophis melanoleucus (Daudin), Nerodia sipedon (Linnaeus), Regina grahami Baird and Girard, Storeria cf. S. dekayi, Thamnophis proximus (Say), Thamnophis radix (Baird and Girard), and Crotalinae indet were recovered.
1989 Platt, Dwight R. Seasonal activity of snakes on a sand prairie. Pages 251-254 in Proceedings of the 11th North American Prairie Conference University of Nebraska Press, Lincoln. pp.
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1989 Brunson, Ken. The rubber snake award. Kansas Herpetological Society Newsletter (77):19
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21
1990 Kaufman, Glennis A. Population ecology, social organization, and mating systems in the Deer Mouse (Peromyscus maniculatus bairdii) in mixed-grass prairie in Kansas. Dissertation. Kansas State University, Manhattan, Kansas. 187pp.
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Sweet, Samuel S. and William S. Parker. Pituophis melanoleucus. Catalogue of American Amphibians and Reptiles (474):1-8
1990 Joy, Jack. An additional note on Howard K. Gloyd. Bulletin of the Chicago Herpetological Society 25(10):180
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Collins, Joseph T. and Suzanne L. Collins. Reptiles and Amphibians of the Cimarron National Grasslands, Morton County, Kansas. U. S. Forest Service, Elkhart, Kansas. 60pp.
1991 Fitch, Henry S. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas. pp.
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1992 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17
1992 Ball, Robert L. High plains serpents: Results of a long-term study in Texas County, Oklahoma and Morton County, Kansas Kansas Herpetological Society Newsletter (88):16-17
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Lokke, John L. Some thoughts on the status of the Western Fox Snake (Elaphe vulpina) in southeast Nebraska, southwest Iowa, northwest Missouri, and northeast Kansas Kansas Herpetological Society Newsletter (89):14-16
1992 Edds, David R. Observations of the 1992 Sharon Springs rattlesnake roundup. Kansas Herpetological Society Newsletter (90):11
1992 Taggart, Travis W. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4
1992 Taggart, Travis W. KHS field trips. Kansas Herpetological Society Newsletter (91):3
1993 Fitzgerald, Eve C. and Charles H. Nilon. Testing the accuracy of an HSI model in and urban county. ():16
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1993 Viets, Brian E. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113
1993 Fitch, Henry S. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224
1994 Fitzgerald, Eve C. Habitat Suitability Index Models for Three Threatened Snake Species in an Urban County. Thesis. University of Missouri, Columbia. pp.
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1994 Riedle, J. Daren. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13
1994 Dloogatch , Michael A. (Editor) Herpetology 1994 Bulletin of the Chicago Herpetological Society 29(12):282-283
Note on the Fitch (1993) paper in the Transactions of the Kansas Academy of Sciences (96(3-4): 213-224) on the abundance.
1995 Holman, J. Alan. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243pp.
1995 Anderson, Lewis, Mark Shaw, Jeff Blodig, and Tom Walker. Report to the Kansas Department of Wildlife and Parks: Herps encountered during REmap project, summer 1994. Kansas Herpetological Society Newsletter (99):10-17
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Parmley, Dennis and J. Alan Holman. Hemphillian (Late Miocene) snakes from Nebraska, with comments on Arikareean through Blancan Snakes of midcontinental North America. Journal of Vertebrate Paleontology 15(1):79-95
1995 Reichling, Steven B. The taxonomic status of the Louisiana pine snake (Pituophis melanoleucus ruthveni) and its relavance to the evolutionary species concept. Journal of Herpetology 29(2):186-198
1996 Miller, Larry L. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Rundquist, Eric M. Notes on the natural history of some Kansas amphibians and reptiles: Parasites. Kansas Herpetological Society Newsletter (105):16-17
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1996 Wright, J. Variations within populations of the Bullsnake. Reptile & Amphibian Magazine (September-October):16-22
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1997 Rundquist, Eric M. Bullsnake combat in Sedgwick County, Kansas. Kansas Herpetological Society Newsletter (108):19
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1998 Gamble, Jerre. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Riedle, J. Daren. Winter snake activity. Kansas Herpetological Society Newsletter (111):16
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1998 Collins, Joseph T. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
1999 Taggart, Travis W. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6
Reported Anaxyrus woodhousii was likely A. fowleri.
2000 Fitch, Henry S. Population structure and biomass of some common snakes in central North America. Scientific Papers of the Natural History Museum University of Kansas (17):1-7
2000 Taggart, Travis W. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5
2000 Van Doren, Mark D. and Curtis J. Schmidt. A herpetological survey of the Fort Larned National Historic Site, Pawnee County, Kansas. Kansas Herpetological Society Newsletter (120):8-11
2000 Taggart, Travis W. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2000 Rodriguez-Robles, Javier A. and Jose M. De Jesus-Escobar. Molecular systematics of New World Gopher, Bull, and Pinesnakes (Pituophis: Colubridae), a transcontinental species complex. Molecular Phylogenetics and Evolution 14(1):35–50
2001 Johnson, Richard W. Spatial ecology of the Eastern Coachwhip (Masticophis flagellum flagellum) in and eastern Texas upland community. Thesis. Stephen F. Austin State University, Nacogdoches, Texas. 54pp.
2001 Taggart, Travis W. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14
2001 Schmidt, Curtis J. The amphibians, turtles, and reptiles of the Smoky Valley Ranch, Logan County, Kansas. Kansas Herpetological Society Newsletter (124):9-11
2001 Taggart, Travis W. Results of the KHS spring field trip west. Kansas Herpetological Society Newsletter (125):10
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2001 Kretzer, Justin E. and Jack F. Cully, Jr. Effects of Blacktailed Prairie Dogs on reptiles and amphibians in Kansas shortgrass prairie. Southwestern Naturalist 46(2):171-177
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Fogell, Daniel D. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6pp.
2002 Riedle, J. Daren and A. Hynek. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20
2002 Ellis, Mark R. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5
2002 Taggart, Travis W. Results of the spring 2002 KHS field trip. Journal of Kansas Herpetology (3):6-7
To the Cimarron National Grassland, in Morton and Stevens counties.
2002 Taggart, Travis W. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13
To Washington County, Kansas (also includes ancillary counts from Clay and Marshall counties).
2003 Pitman, James C. Lesser Prairie-Chicken nest site selection and nest success, juvenile gender determination and growth, and juvenile survival and dispersal in southwestern Kansas. Thesis. Kansas State University, Manhattan. pp.
2003 Fogell, Daniel D. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59pp.
This is the version the author submitted to the NPS. Their final publication was modified.
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Taggart, Travis W. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12
2003 Platt, Dwight R. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20
2003 Fitch, Henry S. Reproduction in snakes of the Fitch Natural History Reservation in northeastern Kansas. Journal of Kansas Herpetology (6):21-24
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Taggart, Travis W. Logan County herp count. Journal of Kansas Herpetology (7):8
2003 Fitch, Henry S., Scott Sharp, and Kylee Sharp. Snakes of the University of Kansas biotic succession area. Journal of Kansas Herpetology (8):20-21
2003 Setser, Kirk and John F. Cavitt. Effects of burning on snakes in Kansas, USA, tallgrass prairie. Natural Areas Journal 23(4):315-319
2004 Schmidt, Curtis J. Natural history and status of the exploited Prairie Rattlesnake (Crotalus viridis) in western Kansas and a herpetofaunal inventory of the Smoky Valley Ranch, Logan County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. pp.
2004 Delisle, Jennifer M. and William H. Busby. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Taggart, Travis W. Kansas Herpetological Society 2004 spring field trip. Journal of Kansas Herpetology (9):2
2004 Taggart, Travis W. Results of the 2004 KHS spring field trip to Logan County. Journal of Kansas Herpetology (10):2-7
2004 Washburne, M. Ellsworth County herp count. Journal of Kansas Herpetology (11):10
2004 Collins, Suzanne L. Geographic distribution. Pituophis catenifer. Journal of Kansas Herpetology (11):14
County record, Thomas County, Kansas.
2004 Schmidt, Curtis J. Geographic distribution: Pituophis catenifer. Kansas. Journal of Kansas Herpetology (11):14
County record, Linn County, Kansas.
2004 Taggart, Travis W. Results of the KHS 2004 fall feld trip . Journal of Kansas Herpetology (12):15-16
2005 Taggart, Travis W. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21
2005 Pitman, James C., Christian A. Hagen, Brent E. Jamison, Robert J. Robel, Thomas M. Loughin, and Roger D. Applegate. Location and success of Lesser Prairie-Chicken nests in relation to vegetation and human disturbance. Journal of Wildlife Management 69(3):1259-1269
2006 Taggart, Travis W. Addendum report to biological inventory of the sandsage prairie near Holcomb, Kansas. Sunflower Electric Cooperative, Hays, Kansas. 31pp.
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Fitch, Henry Sheldon. Gopher snakes, bullsnakes and pine snakes. Journal of Herpetology (17):16-17
2006 Wilgers, Dustin J. and Eva A. Horne. Effects of different burn regimes on tallgrass prairie herpetofaunal species diversity and community composition in the Flint Hills, Kansas. Journal of Herpetology 40():73-84
2006 Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62():378-388
2006 Pitman, James C., Christian A. Hagen, Brent E. Jamison, Robert J. Robel, Thomas M. Loughin, and Roger D. Applegate. Nesting ecology of Lesser Prairie Chickens in Sand Sagebrush Prairie of southwestern Kansas. The Wilson Journal of Ornithology 118(1):23-35
2007 Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5pp.
2007 Kapfer, Joshua M. Ecology of Bullsnakes (Pituophis catenifer sayi) in the upper midwest. Dissertation. University of Wisconsin-Milwaukee, Milwaukee, Wisconsin. 79pp.
2007 Taggart, Travis W. A biological inventory of the Sunflower Electric Site near Holcomb, Kansas. Journal of Kansas Herpetology 23():11-16
2007 Hagen, Christian A., James C. Pitman, Brett K. Sandercock, Robert J. Robel, and Roger D. Applegate. Age-specific survival and probable causes of mortality in female Lesser Prairie Chickens. Journal of Wildlife Management 71(2):518-525
2008 Taggart, Travis W. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3
2008 Iverson, John B; Cameron A. Young, and Thomas S. Akre. Body size and growth in the Bullsnake (Pituophis catenifer sayi) in the Nebraska Sandhills Journal of Herpetology 42(3):501-507
2008 Joshua M. Kapfer, James R. Coggins, and Robert Hay Spatial ecology and habitat selection of Bullsnakes (Pituophis catenifer sayi) at the northern periphery of their geographic range. Copeia 2008(4):815-82
2010 Miller, Larry L. Investigation of the Checkered Garter Snake in Kansas with notes on other Amphibians, Reptiles, and Turtles encountered. Kansas Department of Wildlife and Parks, Pratt, Kansas. 31pp.
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2011 Houck, Mike. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2012 LeClere, Jeffrey B., Erica P. Hoaglund, Jim Scharosch, Christopher E. Smith, Tony Gamble. Two naturally occurring intergeneric hybrid snakes (Pituophis catenifer sayi × Pantherophis vulpinus; Lampropeltini, Squamata) from the midwestern United States Journal of Herpetology 46(2):257-262
2012 Iverson, John B., Cameron A. Young, Thomas S. B. Akre, and Christopher M. Griffiths Reproduction by female bullsnakes (Pituophis catenifer sayi) in the Nebraska Sandhills. The Southwestern Naturalist 57(1):58-73
2012 Martino, Jessica A., Ray G. Poulin, Dennilyn L. Parker, and Christopher M. Somers. Habitat selection by grassland snakes at northern range limits: Implications for conservation. The Journal of Wildlife Management 76(4):759-767
2013 Kauffman, Greg Lee. Stable isotope analysis of a middle woodland population from north central Kansas. Thesis. University of Kansas, Lawrence, Kansas. 110pp.
2013 Sinclair, Tom. A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9
2013 Baldwin, Mary Kate. “Herps in Havensville” Herp Surveys. Collinsorum 2(3/4):10
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2012 Summer Field Trip to Meade County State Park. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Summer Field Trip to Coldwater Lake, Comanche County. Collinsorum 2(3/4):5
2013 Taggart, Travis W. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2013 Gardiner, Laura E., Christopher M. Somers, Jessica A. Martino, Dennilyn L. Parker, and Ray G. Poulin. Balancing the dumbbell: Summer habitats need protection in addition to winter dens for northern snake communities. The Journal of Wildlife Management 77(5):975-982
2014 Helms, Clinton. Nest Survival of Grassland Breeding Birds in a Southern Mixed-Grass Prairie Wetland. Thesis. Fort Hays State University, Hays, Kansas. 100pp.
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2014 Taggart, Travis W. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12
2014 Taggart, Travis W. Results of the 2014 KHS summer field trip to Morton County and adjacent Colorado, New Mexico, and Oklahoma. Collinsorum 3(2-4):12
2014 Taggart, Travis W. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10
2015 Brown, Kasandra A. Occupancy Modeling Of Herpetofauna And Grassland Nesting Birds At Quivira National Wildlife Refuge. Thesis. Fort Hays State University, Hays, Kansas. 72pp.
2015 Mike Houck. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2015 Taggart, Travis W. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3
2016 Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. Vertebrate animals on the Fitch Natural History Reservation (1948-2002) Kansas Biological Survey Report Number 188, Lawrence. 48pp.
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):102
2017 Hutto, Paxson and Travis W. Taggart. Recent sightings of the Western Diamond-backed Rattlesnake at Kanopolis State Park, Ellsworth County, Kansas. Collinsorum 6(1):13
2017 Mardis, Dexter R. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10
2017 Taggart, Travis W. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8
2018 Houck, Mike. Herp Count: Fort Riley Military Installation Collinsorum 7(1):17
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2019 Riedle, J. Daren. The truth about snakes. Kansas Wildlife and Parks Magazine July/August():18-21
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2021 Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. Herp Count: Pratt County: KHS-2020-05 Collinsorum 9(3):12
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Hamilton County: KHS-2020-10. Collinsorum 9(3):13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Morton County: KHS-2020-12. Collinsorum 9(3):13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Seward County: KHS-2020-13. Collinsorum 9(3):13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Stanton County: KHS-2020-10. Collinsorum 9(3):13
2021 Falcon, Melanie. Herp Count: McPherson County: KHS-2020-17. Collinsorum 9(3):14
2021 Taggart, Travis W. Herp Count: Russell County: KHS-2020-24. Collinsorum 9(3):14-15
2021 Taggart, Megan M. and Travis W. Taggart. Herp Count: Seward County: KHS-2020-27. Collinsorum 9(3):15
2021 Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. Herp Count: Stafford County: KHS-2020-34. Collinsorum 9(3):16
2021 Bezy, Robert L. Thoughts on Gophersnakes. Sonoran Herpetologist 34(4):111-122
2023 Russell, Elisabeth Habitat associations and fine-scale movements of the Red-spotted Toad (Anaxyrus punctatus) in Kansas and the efficacy of remote telemetry for monitoring small-scale movements. Thesis. Fort Hays STate University, Hays, Kansas. 81pp.
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Travis W. Taggart © 1999-2024 — w/ Sternberg Museum of Natural History, Fort Hays State University