An adult Plains Gartersnake from Wichita County. © Travis W. Taggart.
An adult Plains Gartersnake from Rush County. Image © Jill Wussow.
An adult Plains Gartersnake from Meade County. Image © Travis W. Taggart.
An adult Plains Gartersnake from Ellis County, Kansas. © Suzanne L. Collins, CNAH.
A sub-adult Plains Gartersnake from Pratt County. © Daren Riedle.
An adult Plains Gartersnake from Logan County. Image © Angie Riesch.
REPTILIA (Reptiles) SQUAMATA (PART) (Snakes) NATRICIDAE (Harmless Egg-Retaining Snakes)

Plains Gartersnake
Thamnophis radix (Baird & Girard 1853)
thăm-nō-fĭs — rā-dĭx


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
HARMLESS. The Plains Gartersnake is characterized by keeled scales, a single anal scale, a stripe on each side of the body situated on the third and fourth scale rows (counting from the belly up), dark vertical bars on the upper lips, and alternating rows of black spots between the stripes on the back. This snake is greenish gray, light olive, or tan, with black spots between the three stripes on the body. The stripe running down the middle of the back may be bright yellow or orange; the stripes on the sides are normally yellow. The belly is white, grayish, or greenish, with a row of black spots down each side. Some specimens of this snake from Barton County in central Kansas exhibit a bright red color on the sides of their bodies. A color photograph of this variant appears in Behler and King (1979). Adult males have longer tails than females. Females grow slightly larger than males.Young look like miniature adults.
Adults normally grow to 38.0-71.0 mm (15-28 inches) in total length. The largest specimen from Kansas is a female (KU 221508) from Ellis County with a total length of 104.4 mm (41­1⁄8 inches) collected by Mark Van Doren, Curtis J. Schmidt, and Russell Toepfer in September 1993. The maximum length throughout the range is 109.2 cm (43 inches) (Powell et al., 2016).

Distribution:
Found in all but the southeastern corner of the state. Seldom encountered east of the Flint Hills or in the central portion of the Red Hills where populations become localized.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
Open icons are questionable records; Click on a marker to view details.
Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 1,096
    Records 
  • 1,032
    Museum Vouchers 
  • 64
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Atchison (1); Barber (4); Barton (109); Butler (2); Chase (1); Cheyenne (22); Clark (3); Clay (1); Cloud (4); Coffey (7); Comanche (1); Cowley (1); Decatur (6); Dickinson (2); Douglas (55); Edwards (2); Ellis (96); Ellsworth (8); Finney (40); Ford (19); Franklin (26); Geary (2); Gove (12); Graham (3); Grant (6); Gray (6); Greeley (2); Greenwood (2); Hamilton (29); Harper (5); Harvey (9); Haskell (1); Hodgeman (7); Jefferson (2); Jewell (4); Kearney (12); Kearny (1); Kingman (2); Kiowa (4); Lane (10); Leavenworth (1); Lincoln (1); Logan (31); Lyon (11); Marion (6); McPherson (5); Meade (59); Mitchell (2); Morris (2); Morton (49); Ness (3); Norton (2); Osage (1); Osborne (5); Ottawa (1); Pawnee (7); Phillips (1); Pottawatomie (2); Pratt (4); Rawlins (3); Reno (32); Republic (6); Rice (1); Riley (18); Rooks (16); Rush (9); Russell (49); Saline (5); Scott (11); Sedgwick (17); Seward (5); Shawnee (1); Sheridan (5); Sherman (18); Stafford (49); Stanton (6); Stevens (1); Sumner (1); Thomas (3); Trego (27); Unknown (72); Wabaunsee (1); Wallace (16); Wichita (2);

Fossil History:
Pleistocene fossil specimens are known from Meade, Rice, and Jewell counties.
Fossils from the Williams Local Fauna of Rice County (Pleistocene: Illinoian) (Preston 1979; Holman 1984; Holman 1995) are assignable to this taxon.
Fossils from the Mount Scott Local Fauna of Meade County (Pleistocene: Illinoian) (Preston 1979, Holman 1987; Holman 1995) are assignable to this taxon.
Fossils from the Butler Spring Local Fauna of Meade County (Pleistocene: Illinoian) (Tihen 1962; Preston 1979; Holman 1995) are assignable to this taxon.
Fossils from the Cragin Quarry Local Fauna of Meade County (Pleistocene: Sangamonian) (Hay 1917; Etheridge 1958; Etheridge 1960, Tihen 1960; Tihen 1962; Brattstrom 1967; Preston 1979; Holman 1995) are assignable to this taxon (species indeterminate).
Fossils from the Courtland Canal local fauna of Jewell County (Eshelman and Hager, 1984) are referable to this taxon.

Natural History:
Plains Gartersnakes prefer open grassy prairies, particularly along the edges of streams, marshes, and lakes. They are normally active from March to November, but during a warm spell in the coldest winter months they will emerge from their underground retreats. This snake is usually active during the day, basking in the sun or foraging for food. Berglund (1967) noted that at Cheyenne Bottoms, Barton County, this species is killed frequently by automobiles because it commonly basks in the sun on dirt or gravel roads.
Knight and Collins (1977) observed that Plains Gartersnakes were found exclusively along streams in Cheyenne County during late May. Platt (1985) studied the habitat preference of this snake in Harvey County and found it abundant in sand prairies. Irwin and Collins (1987) found 25 of these serpents along the stillwater ditches of Cheyenne Bottoms from 1 May to 6 June.
This snake mates during April and May and sometimes in the fall. One or more courting males crawl over and alongside a female with jerking, writhing movements. A successful male curls his tail beneath the female's until their cloacal openings meet and copulation occurs. More than one male may mate with a female. During late July, August, or September, each female gives birth to a litter of young which may range in number from five to sixty (Fitch, 1985), with an average of twenty. Gloyd (1928) reported a female from Franklin County which gave birth to thirteen young on 6 August. Collins (1974) obtained a female on 15 June in Ellsworth County which gave birth to twenty young on 18 September. Eric M. Rundquist (pers. comm., 1989) recorded a litter of ten young born on 30 July to a female from Sumner County.
The Plains Gartersnake feeds on earthworms, toads, frogs, salamanders, fishes, and small rodents. Gray and Douglas (1989) recorded this snake preying on Southern Redbelly Dace in isolated pools on the Konza Prairie near Manhattan.
Predators of the Plains Gartersnake include large birds, mammals, and other snakes (Collins, 1993).

Occurrence Activity:
Number of Unique Obervations (=days): 212; Range: 02 Jan to 30 Dec
Remarks:
The Plains Gartersnake was first reported from Kansas by Mozely (1877). The first specimen (USNM 329) was collected near the "Little Blue River" on 29 July 1857.
The records for Coffey, Osage, and Franklin counties are in need of corroboration. Any reports of specimens east of the Flint Hills are noteworthy.
A series from Coffey County (KU 1997, 2028, 2031, 2044-7) were collected on 27 June 1912, but their given locality "Neosho River" is too ambiguous to plot.
Rundquist (1999) reported assumed hybridization with a captive female Thamnophis sirtalis, however, given that long-term sperm storage and multiple paternity have been reported in T. sirtalis, there are other interpretations not considered.
Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity for this snake of eight years, five months, and three days.

Bibliography:
1863 Jan, Giorgio. Elenco Sistematico degli Ofidi descriti e disegnati per l'Iconografia Generale. [Systematic list of Ophidians described and drawn for General Iconography.] A. Lombardi, Milan, Italy. 143pp.
Records Heterodon nasicus, Lampropeltis calligaster, Coluber constrictor, and Thamnophis radix from Kansas. The Crotalus viridis (confluentus [sic]) record is likely a Sistrurus tergeminus (though S. tergeminus records were also listed separately). It is from Pole Creek, Kansas. is likely (Pole Creek, Washington County, Kansas).
1877 Mozley, Annie E. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6():34-35
First attempt to compile a complete list of snakes reported from Kansas. Most specimens were from Douglas and Wallace counties.
Contains obvious identification/locality errors Thamnophis marcianus (Douglas County) and Heterodon simus (Wallace County).
Neroida holbrookii = Nerodia rhombifer.
Use of the name 'Kansas State University' actually refers to The University of Kansas where Annie E. (Mozley)Boddington received her BS @ KU in 1878 (see also Gloyd 1928).
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():112-123
Also listed the Scarlet Snake (Cemophora coccinea) [=Ophibolus doliatus var. coccineus] and Red Salamander (Pseudotriton ruber) [=Spelerpes ruber] from Kansas.
1882 Brons, H. A. Notes on the habits of some western snakes. The American Naturalist 16(7):564-567
1903 Branson, Edwin B. Snakes of Kansas Thesis. University of Kansas, Lawrence. 41pp.
Describes all snake species reported from Kansas, considering doubtful those species that the author has not encountered himself. This list contains twenty-nine valid species. Collections examined included State University (KU), State House, Washburn College (WU), Ottawa University, State Normal School (ESU), State Agricultural College (KSU), and several high schools in Kansas. Examined the material available to Mozley (1878) and determined that of the thirty-three species listed, only twenty-three species were valid currently. And that Cragin's (1880) list of thirty-two species included eight specimens on the authority of Mozley that this author could not verify in any collection.
1904 Branson, Edwin B. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430
1908 Ruthven, Alexander G. Variations and genetic relationships of the garter-snakes. Bulletin of the United States National Museum (61):1-201
1911 Hurter, Julius. Herpetology of Missouri. Transactions of the Academy of Science St. Louis 20(5):59-274
1927 Burt, Charles E. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12
Accounts on forty species known from the vicinity of Manhattan, Riley County, Kansas. There have been several scientific names changes since publication, which is understandable... however, some usages cannot be explained by subsequent taxonomic reappraisals(e.g.  Gastrophryne carolinensis for Gastrophryne olivacea). The writer lists Eumeces (=Plestiodon) obsoletus and E. guttulatus yet correctly reasoning that the latter is the young of the former. Within the account of Tantilla gracilis (a common form) the writer mentions that T. nigriceps was reported from Riley County by Branson (1904) but that the specimens at Kansas State were absent at the time of publication. The Prairie Rattlesnake (Crotalus viridis) was first reported from Riley County by Branson (1904) and is reported by the writer based on a specimen in the museum at Kansas State University by Professor E. A. Popenoe. The Prairie Rattlesnake is not currently native to Riley County, and closest reliable localities are 150 to the west.
1929 Taylor, Edward H. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62
1929 Taylor, Edward H. List of reptiles and batrachians of Morton County, Kansas, reporting species new to the state fauna. University of Kansas Science Bulletin 19(6):63-65
Annotated listing of 25 species discovered in southwest Morton County just prior to the Dust Bowl that wreaked havoc on the region and the subsequent creation of the Cimarron National Grassland. Of special note are Heterodon platirhinos, Thamnophis marcianus, and Anaxyrus debilis (all of which have not been found in the area since).
1933 Stejneger, Leonhard and Thomas Barbour. A Checklist of North American Amphibians and Reptiles. 3rd Edition. Harvard University Press, Cambridge, Massachusetts. pp.
Reference to Kansas is the listed range of several species.
1933 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1935 Burt, Charles E. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16(3):311-366
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1937 Grant, Chapman. Herpetological notes from Central Kansas. American Midland Naturalist 18(3):370-372
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1938 Schmidt, Karl P. Herpetological evidence for the postglacial eastward extension of the steppe in North America. Ecology 19(3):396-407
1939 Tihen, Joseph A. and James M. Sprague. Amphibians, reptiles, and mammals of the Meade County State Park Transactions of the Kansas Academy of Science 42():499-512
1941 Schmidt, Karl Peterson and D. D. Davis. Field Book of Snakes of the United States and Canada. C.P. Putnam and Sons, New York. 365pp.
1942 Hudson, G. E. The amphibians and reptiles of Nebraska. Nebraska Conservation Bulletin 24():1-146
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1949 Smith, Albert G. The subspecies of the Plains garter snake, Thamnophis radix. Bulletin of the Chicago Academy of Sciences 8(14):285-300
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1951 Wolfenbarger, Keith. A. Systematic and Biological Studies on North American Chiggers of the genus Eutrombicula (Acarina, Trombiculidae). Thesis. University of Kansas, Lawrence. 77pp.
1951 Brumwell, Malcolm J. An ecological survey of the Fort Leavenworth Military Reservation American Midland Naturalist 45(1):187-231
Published posthumously. Lieutenant Brumwell died December 14, 1941, as a result of injuries incurred during the Japanese attack on Pearl Harbor. This paper is a condensed version of his thesis for the Master's degree.
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1960 Steward, P. L. Lung-flukes of snakes, genera Thamnophis and Coluber, in Kansas. University of Kansas Science Bulletin 41():877-890
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1967 Choate, Jerry R. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46pp.
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1975 Rundquist, Eric M. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3pp.
Short accounts for twenty-nine recognized amphibians and reptiles from Kingman County, Kansas. With habitat descriptions and for some species, estimates of population density.
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Capron, Marty B. and Jan Perry. A July weekend in Great Bend. Kansas Herpetological Society Newsletter (14):1-2
1977 Knight, James L. and Joseph T. Collins. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19pp.
1977 Lesch, Martin E. Autecology of the Western Plains Garter Snake, Thamnophis radix haydeni (Reptilla[sic]: Colubridae) in Douglas County, Nebraska. Thesis. University of Nebraska at Omaha, Omaha, Nebraska. 51pp.
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
Notable mentions: Spotted Salamander, Smooth Green Snake
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1979 Ports, Mark A. Occurrence and density studies of nongame wildlife in southwestern Kansas - May 16-August 16, 1979. Kansas Fish and Game Commission, Pratt, Kansas. 83pp.
1979 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1978. Technical Publication of the State Biological Survery of Kansas 8():56-66
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilisThamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1980 Spencer, Dwight. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64pp.
1980 Gray, Peter. An albino Thamnophis radix haydeni (Western Plains Garter Snake). Herpetological Review 11():112
Blurb at the bottom of the page of text accompanied by a b/w photo of the snake. Text reads: An albino THAMNOPHIS RADIX HAYDENI (Western Plains Garter Snake). USA. KANSAS: Sedgwick Co., 9.6 km N, 3.2 km W Clearwater on dirt road. 18 April 1978. T. L. Sullivan. In the collection of the Sedgwick County Zoological Society. Photograph by P. Gray and J. Marlett. Pentax with Tri-X at f 7.0 at 500th sec. Contributed by P. Gray.
1980 Glass, G. E., and N .A. Slade. The effect of Sigmodon hispidus on spatial and temporal activity of Microtus ochrogaster: Evidence for competition Ecology 61():358-370
1981 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survery of Kansas 10():7-19
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1982 Rogers, Karel L. Herpetofaunas of the Courland Canal and Hall Ash Local Faunas (Pleistoncene: Early Kansas) of Jewell Co., Kansas. Journal of Herpetology 16(2):174-177
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1984 Von Achen, Pennie. H. and John L. Rakestraw. The role of chemoreception in the prey selection of neonate reptiles. Pages 163-172 in Vertebrate Ecology and Systematics— A Tribute to Henry S Fitch. Special Publication Number 10. The University of Kansas Museum of Natural History, Lawrence. pp.
1984 Holman, J. Alan. Herpetofaunas of the Duck Creek and Williams Local Faunas (Pleistocene: Illinoian) of Kansas. Pages 20-38 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. pp.
1984 Eshelman, Ralph and Michael Hager. Two Irvingtonian (Medial Pleistocene) vertebrate faunas from northcentral Kansas. Pages 384-404 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8 Special Publication Number 8, Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. pp.
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Heinrich, Mark L. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57pp.
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1984 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26
Invalidated the specimens of Thamnophis sirtalis from Hamilton County (reidentified as T. cyrtopsis; KU 2088) and Wallace County mapped in Collins, 1982.
1985 Lynch, John D. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Science 13():33-57
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1986 Holman, J. Alan. Butler Spring herpetofauna of Kansas (Pleistocene: Illinoian) and its climatic significance. Journal of Herpetology 20(4):568-569
1987 Coleman, Keith. Annual KHS Field Trip held at Atchison State Lake. Kansas Herpetological Society Newsletter (68):5-6
1987 Holman, J. Alan. Climatic significance of a late Illinoian herpetofauna from southwestern Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(5):129-141
Anura - Bufo sp., ?Acris sp., Rana pipiens complex
Testudines - Sternotherus odoratus (Latreille), Chelydra serpentina (Linnaeus), Chrysemys picta (Schneider), Emydoidea blandingii (Holbrook), *Pseudemys hibbardi (Preston), Pseudemys scripta (Schoepff), Terrapene carolina (Linnaeus), Trionyx sp.
Squamata - Ophisaurus attenuatus Baird, Heterodon sp., Diadophis punctatus (Linnaeus), Coluber cf. C. constrictor Linnaeus, Elaphe vulpina (Baird and Girard), Lampropeltis getulus (Linnaeus), Pituophis melanoleucus (Daudin), Nerodia sipedon (Linnaeus), Regina grahami Baird and Girard, Storeria cf. S. dekayi, Thamnophis proximus (Say), Thamnophis radix (Baird and Girard), and Crotalinae indet were recovered.
1987 Holman, J. Alan. Snakes from the Robert Local Fauna (Late Wisconsinan) of Meade County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 27(6):143-150
Heterodon sp. indet, Diadophis punctatus (Linnaeus) Coluber constrictor Linnaeus, Lampropeltis getulus (Linnaeus) Nerodia sipedon (Linnaeus) Regina grahamii, Baird and Girard, Storeria cf. Storeria dekayi (Holbrook), Thamnophis proximus (Say), Thamnophis proxirnus (Say) or Thamnophis sirtalis (Linnaeus), and Thamnophis radix (Baird and Girard) recovered.
1988 Miller, Larry L. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6
1989 Platt, Dwight R. Seasonal activity of snakes on a sand prairie. Pages 251-254 in Proceedings of the 11th North American Prairie Conference University of Nebraska Press, Lincoln. pp.
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1990 Toepfer, Karen. The tail (tail?) of the $1,375.92 garter snake. Kansas Herpetological Society Newsletter (82):12-14
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Collins, Joseph T. and Suzanne L. Collins. Reptiles and Amphibians of the Cimarron National Grasslands, Morton County, Kansas. U. S. Forest Service, Elkhart, Kansas. 60pp.
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1992 Ball, Robert L. High plains serpents: Results of a long-term study in Texas County, Oklahoma and Morton County, Kansas Kansas Herpetological Society Newsletter (88):16-17
1992 Taggart, Travis W. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4
1992 Taggart, Travis W. KHS field trips. Kansas Herpetological Society Newsletter (91):3
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Fitch, Henry S. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224
1994 Taggart, Travis W. The natural history and distribution of the Green Toad (Bufo debilis) in Kansas, with a report on an effort to reintroduce the species into the Cimarron National Grasslands. Kansas Department of Wildlife and Parks, Pratt. 12pp.
1994 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4
See, Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.
1994 Dloogatch , Michael A. (Editor) Herpetology 1994 Bulletin of the Chicago Herpetological Society 29(12):282-283
Note on the Fitch (1993) paper in the Transactions of the Kansas Academy of Sciences (96(3-4): 213-224) on the abundance.
1995 Holman, J. Alan. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243pp.
1995 Anderson, Lewis, Mark Shaw, Jeff Blodig, and Tom Walker. Report to the Kansas Department of Wildlife and Parks: Herps encountered during REmap project, summer 1994. Kansas Herpetological Society Newsletter (99):10-17
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1996 Rossman, Douglas A., Neil B. Ford, and Ricahrd A. Seigel. The Garter Snakes: Evolution and Ecology. University of Oklahoma Press, Norman. pp.
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1997 Taggart, Travis W. Status of Bufo debilis (Anura: Bufonidae) in Kansas Kansas Herpetological Society Newsletter (109):7-12
The specimens (KU) cited from Hamilton County, have been corrected.
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
1999 Rundquist, Eric M. Garter snake hybridization. Kansas Herpetological Society Newsletter (118):15
2000 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1999. Kansas Herpetological Society Newsletter (119):7-9
2000 Taggart, Travis W. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5
2000 Van Doren, Mark D. and Curtis J. Schmidt. A herpetological survey of the Fort Larned National Historic Site, Pawnee County, Kansas. Kansas Herpetological Society Newsletter (120):8-11
2000 Taggart, Travis W. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2000 Murphy, John C. and Russell M. Curry. A case of parthenogenesis in the Plains Garter Snake, Thamnophis radix. Bulletin of the Chicago Herpetological Society 35(2):17-19
2001 Schmidt, Curtis J. The amphibians, turtles, and reptiles of the Smoky Valley Ranch, Logan County, Kansas. Kansas Herpetological Society Newsletter (124):9-11
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Fogell, Daniel D. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6pp.
2002 Taggart, Travis W. Results of the spring 2002 KHS field trip. Journal of Kansas Herpetology (3):6-7
To the Cimarron National Grassland, in Morton and Stevens counties.
2002 Taggart, Travis W. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13
To Washington County, Kansas (also includes ancillary counts from Clay and Marshall counties).
2002 Collins, Suzanne L. Geographic distribution: Thamnophis radix. Journal of Kansas Herpetology (4):14
2003 Fogell, Daniel D. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59pp.
This is the version the author submitted to the NPS. Their final publication was modified.
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 2002. Journal of Kansas Herpetology (5):13-16
2003 Taggart, Travis W. Communal brumaculum. Journal of Kansas Herpetology (5):16
2003 Platt, Dwight R. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20
2003 Burr, Andrew. Coffey County herp count 1. Journal of Kansas Herpetology (7):7
2003 Walley, Harlan D., T. L. Wusterbarth, and Kristin M. Stanford. Thamnophis radix. Catalogue of American Amphibians and Reptiles (779):42748
2004 Schmidt, Curtis J. Natural history and status of the exploited Prairie Rattlesnake (Crotalus viridis) in western Kansas and a herpetofaunal inventory of the Smoky Valley Ranch, Logan County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. pp.
2004 Delisle, Jennifer M. and William H. Busby. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Keathley, Vicki L. Tactile discrimination in three species of Garter Snake (Thamnophis). Dissertation. University of Texas at Arlington, Arlington, Texas. 99pp.
2004 Taggart, Travis W. Kansas Herpetological Society 2004 spring field trip. Journal of Kansas Herpetology (9):2
2004 Taggart, Travis W. Results of the 2004 KHS spring field trip to Logan County. Journal of Kansas Herpetology (10):2-7
2004 Taggart, Travis W. Results of the KHS 2004 fall feld trip . Journal of Kansas Herpetology (12):15-16
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62():378-388
2007 Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5pp.
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2013 Taggart, Travis W. KHS 2012 Summer Field Trip to Meade County State Park. Collinsorum 2(3/4):3
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2014 Taggart, Travis W. Results of the 2014 KHS summer field trip to Morton County and adjacent Colorado, New Mexico, and Oklahoma. Collinsorum 3(2-4):12
2014 Taggart, Travis W. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10
2015 Brown, Kasandra A. Occupancy Modeling Of Herpetofauna And Grassland Nesting Birds At Quivira National Wildlife Refuge. Thesis. Fort Hays State University, Hays, Kansas. 72pp.
2015 Mike Houck. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11
2015 Taggart, Travis W. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):102
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2019 Riedle, J. Daren. The truth about snakes. Kansas Wildlife and Parks Magazine July/August():18-21
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2021 Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. Herp Count: Pratt County: KHS-2020-05 Collinsorum 9(3):12
2021 Riedle, J. Daren. Herp Count: Pratt County: KHS-2020-07 Collinsorum 9(3):12
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Hamilton County: KHS-2020-10. Collinsorum 9(3):13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Hodgeman County: KHS-2020-09. Collinsorum 9(3):13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Morton County: KHS-2020-12. Collinsorum 9(3):13
2021 Taggart, Meg, Amelia Jaeger, Jesse J. Taggart, and Travis W. Taggart. Herp Count: Russell County: KHS-2020-22. Collinsorum 9(3):14-15
2021 Taggart, Megan M. and Travis W. Taggart. Herp Count: Seward County: KHS-2020-27. Collinsorum 9(3):15
2021 Riedle, J. Daren, Tamera D. Riedle, Zachary Riedle, and Greya Riedle. Herp Count: Stafford County: KHS-2020-34. Collinsorum 9(3):16
2021 Riedle, J. Daren. Herp Count: Barton County: KHS-2020-28. Collinsorum 9(3):16
2021 Schmidt, Curtis J. Herp Count: Ellis County: KHS-2020-32. Collinsorum 9(3):16
2022 Hallas, Joshua M., Thomas L. Parchman, and Chris R. Feldman. Phylogenomic analyses resolve relationships among garter snakes (Thamnophis: Natricinae: Colubridae) and elucidate biogeographic history and morphological evolution. Molecular Phylogenetics and Evolution 167():13
Utilized a genome-wide high-throughput sequencing data and near-complete taxon sampling to estimate a robust phylogeny for Thamnophis.
2023 Nunez, Leroy, Levi N Gray, David Weisrock, and Frank T Burbrink. The phylogenomic and biogeographic history of the Gartersnakes, Watersnakes, and Allies (Natricidae: Thamnophiini). Molecular Phylogenetics and Evolution 186(5:107844):12
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Travis W. Taggart © 1999-2024 — w/ Sternberg Museum of Natural History, Fort Hays State University