NORTH AMERICAN RACER
Coluber constrictor Linnaeus 1758
kō-lū-bĕr — kŏn-strĭk-tŏr


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None

An adult North American Racer from Shawnee County, Kansas. © Lisa Wehrly.
An adult North American Racer from Chautauqua County, Kansas. Image © Dan Fogell.
A juvenile NA Racer from Hodgeman County. © Travis W. Taggart.
An adult North American Racer from Riley County. © Maci Loughrea
An juvenile North American Racer from Logan County. © Travis W. Taggart.
An adult North American Racer from Johnson County. © Nick Edge.
A juvenile from Russell County. Image © Jim Scharosch.
An adult from Ellsworth County. Image © Suzanne L. Collins, CNAH.
An adult North American Racer from Johnson County.  © Nick Edge

Description:
Harmless. The North American Racer has smooth scales. Adults have a uniform blue-gray, greenish-blue, or rarely brown body and a uniform yellow to cream-colored belly with no pattern. Young individuals, have a series of large light-edged blotches down their backs alternating with smaller spots on their sides and scattered dark speckles on the belly.
As North American Racers grow older their pattern is lost as they acquire the uniform appearance of adults. Adult males have slightly longer tails than females, but females grow slightly larger than males.
Snake eggs can be especially difficult to distinguish from one species to another, however, the North American Racer is the only species in Kansas that has roughly textured eggs (like large-grit sandpaper). The eggs are white to cream and typically do not adhere to one another. They are approximately an inch long and one-half inch in diameter.
Adults are normally 580-1,270 mm (23-50 inches) in TL; largest specimen from Kansas: female (KU 192239) from Jefferson County with TL of 1,413 mm (55½ inches) collected by Rick Strawn on 31 May 1982; maximum length throughout the range: 75 inches (Boundy, 1995); maximum weight for Kansas specimen: 538 grams (1 pound, 3 ounces).

Distribution:
A true generalist, this species is found statewide. The North American Racer is a snake of open grassland, pasture, and prairie areas during the summer and generally is found on rocky wooded hillsides only in spring and fall. This snake is active normally from April to mid-November, at air temperature ranging from 60 to 90 F, but Clarke (1958) found it active as early as 23 March in Osage County. Platt (1985) found this reptile common in the sand prairie habitat of western Harvey County.
(, Museum Voucher) (, Observation) (, Literature Record)
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  • Occurrence Summary:  
  • 1,520
    Records 
  • 1,212
    Museum Vouchers 
  • 308
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (12); Anderson (23); Atchison (11); Barber (26); Barton (6); Bourbon (37); Brown (2); Butler (8); Chase (24); Chautauqua (8); Cherokee (35); Cheyenne (11); Clark (10); Clay (2); Cloud (4); Coffey (5); Comanche (15); Cowley (76); Crawford (16); Decatur (9); Dickinson (5); Doniphan (11); Douglas (113); Edwards (3); Elk (16); Ellis (128); Ellsworth (16); Finney (17); Ford (3); Franklin (16); Geary (11); Gove (23); Graham (9); Grant (5); Gray (4); Greeley (1); Greenwood (31); Hamilton (11); Harper (21); Harvey (9); Haskell (3); Hodgeman (11); Jackson (4); Jefferson (23); Jewell (7); Johnson (17); Kearney (5); Kingman (4); Kiowa (13); Labette (4); Lane (4); Leavenworth (15); Lincoln (6); Linn (23); Logan (12); Lyon (37); Marion (3); Marshall (25); McPherson (2); Meade (23); Miami (35); Mitchell (2); Montgomery (5); Morris (2); Morton (8); Nemaha (1); Neosho (9); Ness (12); Norton (3); Osage (5); Osborne (7); Ottawa (2); Pawnee (4); Phillips (10); Pottawatomie (31); Pratt (4); Rawlins (9); Reno (3); Republic (7); Rice (6); Riley (71); Rooks (18); Rush (5); Russell (51); Saline (13); Scott (6); Sedgwick (2); Seward (14); Shawnee (4); Sheridan (1); Sherman (2); Smith (1); Stafford (13); Stanton (6); Stevens (1); Sumner (4); Thomas (5); Trego (24); Unknown (20); Wabaunsee (15); Wallace (15); Washington (7); Wichita (4); Wilson (5); Woodson (3); Wyandotte (9);

Natural History:
On warm days, the North American Racer is often seen gliding over the ground in search of food. Their average home range is 25 acres but they are not territorial. They pursue and eat insects, frogs, lizards, other snakes (including venomous species), birds, bird eggs, and small mammals. Despite its scientific name, it is not a constrictor.
Gloyd (1925) removed a Spotted Ground Squirrel (Xerospermophilus spilosoma) and a 19 inch North American Racer (Coluber constrictor) from the stomach of a Red-tailed Hawk (Buteo jamaicensis) in Kansas in 1921.
Burt (1935) reported that Cornelius Rogers placed a small Prairie Kingsnake (Lampropeltis calligaster} in a cage with an adult North American Racer on 31 August 1934 and the Coluber immediately attacked the smaller snake, swallowing it, tail first.
Breeding in this species occurs in May. During courtship the male positions himself alongside the female and ripples his body spasmodically as he positions his cloaca beneath her tail. Copulation then occurs and lasts several minutes. While copulating, the female may move forward slowly, dragging the attached male with her.
Five to 31 rough-shelled eggs per clutch, with an average of eleven or twelve (Fitch, 1985), are laid in June or July, usually in tunnels or burrows of small mammals, such as moles and often under rocks or slabs of concrete. Incubation takes 2–3 months. More than one female may use the same nest site at the same time. Guillette and Sullivan (1983) reported a case of successful incubation of eggs removed from a pregnant road-killed Racer in Harvey County.
Cope (1900) reports an account, given by Dr. Henry Brons, of the courtship of this species on the Kansas plains: "The manner of union of the sexes at this season is rather instructive. The female among the racers (Bascanium) is larger and darker than the males, and not so graceful in form or movements. She, at times, seems to toy with the male, indisposed to yield to his importunities, though pressed with ardor. To avoid his suit, at times, she will dart through grass, among stones, or enter a crevice. Should he be able to reacher his mate while within a hole, he is not slow in bringing her to the surface, again to be repulsed. Upon an unbroken ground, the sexual communion is less prolonged. Here she is unable to free herself from his quick and effectively directed moves. In case she attempts to quit him, a coil is thrown about her body, and his head laid flat upon her neck and replaced as promptly as dislodged, evidently in the endeavor to propitiate her."
Rundquist (1998) reported on a letter from Claude Hibbard to Edward H. Taylor that recounted the discovery of a North American Racer active on 31 December 1950 at 70 deg F.

Occurrence Activity:
White dates indicate there is at least a single recorded occurrence on that date. The darker blue a date is, the greater the relative number of observations for that date.
Observation Type: (of recorded types)
Remarks:
First reported in Kansas by Hallowell (1856). The earliest existing specimen is from 1871 (MCZ 2456).
Burt (1935) reported on a female North American Racer that was found dead in the road with eggs exposed from the body cavity in Pawnee County, Kansas, on 26 May 1934. Burt (op. cit.) also reported finding young individuals with the initial color pattern in a snake den in Cowley County, Kansas, on 29 March 1934, in association with Pantherophis emoryi and Diadophis punctatus. Adults were associated with large Pitouphis catenifer in another den on a prairie hillside.

Bibliography:
1758 Linné, Carl von (=Linneaus). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis 10th Edition, Volume 1, L. Salvius, Stockholm. iv + 826pp.
1822 Say, Thomas. In James, Edwin. Pages 503 in Account of an expedition from Pittsburgh to the Rocky Mountains, performed in the years 1819, '20, by order of the Hon. J. C. Calhoun, Secretary of War, under the command of Major Stephen H. Long. Vol. 1. H. C. Carey & I. Lea., Philadelphia. pp.
1857 Hallowell, Edward. Note on the collection of reptiles from the neighborhood of San Antonio, Texas, recently presented to the Academy of Natural Sciences by Dr. A. Heerman. Proceedings of the Academy of Natural Sciences, Philadelphia 8():306-310
1857 Hallowell, Edward. Notice of a collection of reptiles from Kansas and Nebraska presented to the Academy of Natural Sciences, by Doctor Hammond, U. S. A. Proceedings of the Academy of Natural Sciences, Philadelphia 8():238-253
1859 Cooper, James G. Report upon the reptiles collected on the survey Pages 292-306 in Cooper, James G. and G. Suckley.  The Natural History of Washington Territory with much relating to Minnesota, Nebraska, Kansas, Oregon, and California. US Government Document, Washington, D. C. pp.
1860 Cooper, J. G. Report upon the reptiles collected on the survey. Pages 292-306 in Cooper, James G. and G. Suckley.  The Natural History of Washington Territory with much relating to Minnesota, Nebraska, Kansas, Oregon, and California. Bailliere Brother, New York, New York. pp.
1860 Cope, Edward D. Supplement to "A catalogue of the venomous serpents in the Museum of the Academy," etc. Supplement to "A catalogue of the venomous serpents in the Museum of the Academy" etc 12():72-74
1877 Mozley, Annie E. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6():34-35
1878 Yarrow, Henry C. and H. W. Henshaw. Report upon the reptiles and batrachians collected during the years of 1875, 1876 and 1877 in California, Arizona and Nevada. Pages 4-23 in Annual Report upon the Geographical Surveys of the Territory of the United States West of the One Hundredth Meridian. Appendix NN , Washington, D. C. pp.
1879 Bocourt, M. F. Etudes sur les reptiles, p. i-xiv, 1-1012. In Recherches Zoologiques pour servir a l'Histoire de Ia Faune de l'Amérique Centrale et du Mexique. Mission Scientifique au Mexique et dans l'Amérique Centrale, Recherches zoologiques. Part 2, sect. 1; In A Imprimerie Impériale, Paris [3, Pt. 6]. A Imprimerie Impériale, Paris, France.. 360–440pp.
1882 Yarrow, Henry C. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249
1886 Ebbutt, Percy G. Emigrant Life in Kansas Swan Sonnenschein and Company, Paternoster Square, London. 237pp.
1900 Cope, Edward D. The crocodilians, lizards and snakes of North America. Pages 153-1270 in Report of the U. S. National Museum for the Year Ending June 30, 1898 , Washington, D. C. pp.
1901 Brown, Arthur Erwin. A review of the genera and species of American snakes, north of Mexico. Proceedings of the Academy of Natural Sciences, Philadelphia 53(1):10-110
1904 Branson, Edwin B. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430
1911 Hurter, Julius. Herpetology of Missouri. Transactions of the Academy of Science St. Louis 20(5):59-274
1917 Wooster, Lyman D. Nature Study Bulletin Kansas State Printing Plant, Topeka, Kansas.. 63pp.
1925 Gloyd, Howard K. Field studies of the diurnal raptores of Eastern and Central Kansas. The Wilson Bulletin 37(3):133-149
1925 Linsdale, Jean M. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312pp.
1927 Linsdale, Jean M. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81
1928 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1929 Taylor, Edward H. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62
1929 Taylor, Edward H. List of reptiles and batrachians of Morton County, Kansas, reporting species new to the state fauna. University of Kansas Science Bulletin 19(6):63-65
1932 Gloyd, Howard K. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Museum of Zoology, University of Michigan 15():389-408
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1935 Burt, Charles E. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16(3):311-366
1935 Burt, Charles E. and William L. Hoyle. Additional records of the reptiles of the central prairie region of the United States Transactions of the Kansas Academy of Science 37():193-216
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1937 Grant, Chapman. Herpetological notes from Central Kansas. American Midland Naturalist 18(3):370-372
1941 Branson, F. and C. Deyoe A study of snakes and lizards of Ellis County. Unpublished data, Fort Hays State University. pp.
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
1955 Auffenberg, Walter. A reconsideration of the racer, Coluber constrictor, in eastern United States. Tulane Studies in Zoology and Botany 2(6):89-155
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1963 Fitch, Henry S. Natural History of the racer Coluber constrictor. University of Kansas Publications Museum of Natural History 15(8):351-468
1970 Fitch, Henry S. Reproductive cycles in lizards and snakes. University of Kansas Museum of Natural History Miscellaneous Publication (52):1-247
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1975 Capron, Marty. A trip through the Kansas Flint Hills. Kansas Herpetological Society Newsletter (8):4-5
1975 Rundquist, Eric M. First KHS field trip yields three county records. Kansas Herpetological Society Newsletter (7):1-3
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1976 Capron, Marty. A trip to the Ozarks. Kansas Herpetological Society Newsletter (15):7-8
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1977 Capron, Marty. An albino Eastern Yellowbellied Racer from Kansas. Kansas Herpetological Society Newsletter (22):7
1977 Knight, James L. and Joseph T. Collins. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19pp.
1977 Perry, Janice. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4
1977 Trott, Gene. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (19):2-3
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
1978 Fitch, Henry S. and E. Raymond Hall. A 20year record of succession on reseeded fields of tallgrass prairie on the Rockefeller Experimental Tract. University of Kansas Museum of Natural History Special Publication (4):1-15
1978 Perry, Janice. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5
1978 Schwaner, Terry D. KHS field trip to Grant County, Kansas, 12-14 May 1978. Kansas Herpetological Society Newsletter (25):3-4
1978 Warner, M. and R. Wencel. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16
1978 Wilson, Larry David. Coluber constrictor. Catalogue of American Amphibians and Reptiles (218):1-4
1979 Gray, Peter and Eddie Stegall. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
1982 Collins, Joseph T. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57pp.
1982 Fitch, Henry S. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C. pp.
1982 Guarisco, Hank. An incidence of albinism in the Eastern Yellowbelly Racer (Coluber constrictor flaviventris). Kansas Herpetological Society Newsletter (49):11
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1983 Guillette, L.J., Jr. and W.P. Sullivan. Life history notes: Coluber constrictor flaviventris. Herpetological Review 14():19
1983 Miller, Larry L. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7
1985 Capron, Marty. Thunder snakes, blow vipers, and others. Kansas Herpetological Society Newsletter (60):9-10
1985 Miller, Larry L. KHS 1985 field trip to Kirwin Reservoir. Kansas Herpetological Society Newsletter (61):11-12
1986 Brunson, Ken. Some unusual injuries to snakes. Kansas Herpetological Society Newsletter (65):13-14
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1986 Schitti, Beat and Larry David Wilson. Coluber. Catalogue of American Amphibians and Reptiles (399):1-4
1987 Simmons, John E. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894
1988 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19
1988 Miller, Larry L. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21
1989 Platt, Dwight R. Seasonal activity of snakes on a sand prairie. Pages 251-254 in Proceedings of the 11th North American Prairie Conference University of Nebraska Press, Lincoln. pp.
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1991 Collins, Joseph T. and Suzanne L. Collins. Reptiles and Amphibians of the Cimarron National Grasslands, Morton County, Kansas. U. S. Forest Service, Elkhart, Kansas. 60pp.
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Fitch, Henry S. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence. pp.
1992 Ball, Robert L. High plains serpents: Results of a long-term study in Texas County, Oklahoma and Morton County, Kansas Kansas Herpetological Society Newsletter (88):16-17
1992 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Edds, David R. Observations of the 1992 Sharon Springs rattlesnake roundup. Kansas Herpetological Society Newsletter (90):11
1992 Rundquist, Eric M. Results of the KHS 1992 fall field trip. Kansas Herpetological Society Newsletter (90):4
1992 Taggart, Travis W. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
1994 Fitzgerald, Eve and Charles Nilon Classification of habitats for endangered and threatened species in Wyandotte County, Kansas Kansas Department of Wildlife and Parks, Pratt, Kansas. 98pp.
1994 Riedle, J. Daren. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13
1994 Riedle, J. Daren. Distribution of the Timber Rattlesnake (Crotalus horridus) in Chautauqua, Elk, and Montgomery counties, Kansas. Kansas Herpetological Society Newsletter (95):43051
1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
1995 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1994. Kansas Herpetological Society Newsletter (100):24-47
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1996 Miller, Larry L. Many amphibian and reptile species identified during KHS 1996 fall field trip to Wabaunsee County. Kansas Herpetological Society Newsletter (106):2-3
1996 Miller, Larry L. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3
1996 Miller, Larry L. Third graders conduct amphibian and reptile field study. Kansas Herpetological Society Newsletter (106):15
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1997 Collins, Joseph T. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3
1997 Rundquist, Eric M. Addendum to 1997 KHS herp counts. Kansas Herpetological Society Newsletter (109):14-15
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1998 Collins, Joseph T. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Gamble, Jerre Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. Racer reproduction and diet observation. Kansas Herpetological Society Newsletter (113):15
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1998 Rundquist, Eric M. Winter snake activity. Kansas Herpetological Society Newsletter (111):16
1998 Simmons, J. E. A technique to remove reptiles from glue boards. Kansas Herpetological Society Newsletter (114):18
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
1999 Taggart, Travis W. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6
2000 Fitch, Henry S. Population structure and biomass of some common snakes in central North America. Scientific Papers of the Natural History Museum University of Kansas (17):1-7
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2000 Taggart, Travis W. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16
2000 Taggart, Travis W. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5
2001 Ellis, Mark R. Reproduction in the Common Garter Snake in Shawnee County, Kansas. Kansas Herpetological Society Newsletter (125):12
2001 Guarisco, Hank. Ode to an ophidion autumn. Kansas Herpetological Society Newsletter (123):19
2001 Kretzer, Justin E. and Jack F. Cully, Jr. Effects of Blacktailed Prairie Dogs on reptiles and amphibians in Kansas shortgrass prairie. Southwestern Naturalist 46(2):171-177
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2001 Schmidt, Curtis J. The amphibians, turtles, and reptiles of the Smoky Valley Ranch, Logan County, Kansas. Kansas Herpetological Society Newsletter (124):9-11
2001 Shewchuk, Christopher H. and James D. Austin. Foot habits of the Racer (Coluber constrictor mormon) in the northern part of its range. Herpetological Journal 11():151-155
2001 Taggart, Travis W. Results of the KHS spring field trip west. Kansas Herpetological Society Newsletter (125):10
2001 Taggart, Travis W. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14
2002 Ellis, Mark R. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Miller, Larry L. Osage County herp count II. Journal of Kansas Herpetology (4):15
2002 Miller, Larry L. Shawnee County herp count. Journal of Kansas Herpetology (4):15
2002 Riedle, J. Daren and A. Hynek. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20
2002 Rundquist, Eric M. Natural history of the Night Snake, Hypsiglena torquata, in Kansas. Journal of Kansas Herpetology (4):16-20
2002 Taggart, Travis W. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13
2002 Taggart, Travis W. Results of the spring 2002 KHS field trip. Journal of Kansas Herpetology (3):6-7
2002 Taggart, Travis W. and Curtis J. Schmidt. Geographic distribution: Coluber constrictor. Journal of Kansas Herpetology (2):10
2003 Burr, Andrew. Coffey County herp count 1. Journal of Kansas Herpetology (7):7
2003 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 2002. Journal of Kansas Herpetology (5):13-16
2003 Fitch, Henry S. Reproduction in snakes of the Fitch Natural History Reservation in northeastern Kansas. Journal of Kansas Herpetology (6):21-24
2003 Fitch, Henry S., Scott Sharp, and Kylee Sharp. Snakes of the University of Kansas biotic succession area. Journal of Kansas Herpetology (8):20-21
2003 Freeman, Craig C. A natural areas inventory of the Ft. Leavenworth Military Reservation, Leavenworth County, Kansas. II. Open-file Report No. 117. Kansas Biological Survey, Lawrence, Kansas. 199pp.
2003 Miller, Larry L. and Suzanne L. Miller. Wakarusa herp count. Journal of Kansas Herpetology (7):10
2003 Platt, Dwight R. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Suleiman, Gibran. Fort Riley herp count. Journal of Kansas Herpetology (7):9
2003 Taggart, Travis W. Communal brumaculum. Journal of Kansas Herpetology (5):16
2003 Taggart, Travis W. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12
2003 Taggart, Travis W. Logan County herp count. Journal of Kansas Herpetology (7):8
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Taggart, Travis W. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15
2003 Volkmann, Al. Cowley County herp count 1. Journal of Kansas Herpetology (7):7
2004 Collins, Joseph T. Marais des Cygnes herp count. Journal of Kansas Herpetology (11):11
2004 Delisle, Jennifer M. and William H. Busby Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Ellis, Mark R. and Kathy Ellis. Wakarusa herp count. Journal of Kansas Herpetology (11):12
2004 Fitch, Henry S. Food surplus and body size in local populations of snakes. Journal of Kansas Herpetology (10):14-16
2004 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (11):11-12
2004 Taggart, Travis W. Kansas Herpetological Society 2004 spring field trip. Journal of Kansas Herpetology (9):2
2004 Taggart, Travis W. Results of the 2004 KHS spring field trip to Logan County. Journal of Kansas Herpetology (10):2-7
2004 Taggart, Travis W. Results of the KHS 2004 fall feld trip . Journal of Kansas Herpetology (12):15-16
2004 Volkmann, Al. Cowley County herp count. Journal of Kansas Herpetology (11):10
2005 Taggart, Travis W. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21
2006 Taggart, Travis W. Addendum report to biological inventory of the sandsage prairie near Holcomb, Kansas. Sunflower Electric Cooperative, Hays, Kansas. 31pp.
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Taggart, Travis W. Results of the KHS spring field trip to Kiowa County. Journal of Kansas Herpetology (18):2-5
2007 Pyron, R. Alexander. A revised distribution record of Masticophis in Kansas. Journal of Kansas Herpetology (24):16
2007 Taggart, Travis W. A biological inventory of the Sunflower Electric Site near Holcomb, Kansas. Journal of Kansas Herpetology 23():11-16
2007 Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5pp.
2008 Taggart, Travis W. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3
2009 Murrow, Daniel G. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
2010 Miller, Larry L. 2010 Investigation of the Checkered Garter Snake in Kansas with notes on other Amphibians, Reptiles, and Turtles encountered. Kansas Department of Wildlife and Parks, Pratt, Kansas. 31pp.
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Houck, Mike. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9
2011 McMartin, D. Chris. Herp Count: Fort Leavenworth Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):8-9
2011 McMartin, David C. U. S. Army 2011 Fort Leavenworth Herpetofaunal Survey: 23 April - 09 May 2011. Privately printed, Leavenworth, Kansas. 33pp.
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2011 Taggart, Travis W. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2013 Baldwin, Mary Kate. “Herps in Havensville” Herp Surveys. Collinsorum 2(3/4):10
2013 Miller, Larry L. Wellington Lake Herpetological Survey. Collinsorum 2(1/2):12
2013 Sinclair, Tom A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9
2013 Taggart, Travis W. KHS 2012 Fall Field Trip to Atchison County State Lake. Collinsorum 2(3/4):4
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2012 Summer Field Trip to Meade County State Park. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6
2013 Taggart, Travis W. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4
2013 Taggart, Travis W. KHS 2013 Summer Field Trip to Coldwater Lake, Comanche County. Collinsorum 2(3/4):5
2014 McMartin, D. Chris. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10
2014 Taggart, Travis W. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10
2014 Taggart, Travis W. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2014 Taggart, Travis W. Results of the 2014 KHS summer field trip to Morton County and adjacent Colorado, New Mexico, and Oklahoma. Collinsorum 3(2-4):12
2015 Brown, Kasandra A. Occupancy Modeling Of Herpetofauna And Grassland Nesting Birds At Quivira National Wildlife Refuge. Thesis. Fort Hays State University, Hays, Kansas. 72pp.
2015 Mike Houck. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11
2015 Taggart, Travis W. Fall Field Trip Held In Washington County. Collinsorum 4(3):4
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2015 Taggart, Travis W. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3
2016 Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle Vertebrate animals on the Fitch Natural History Reservation (1948-2002) Kansas Biological Survey Report Number 188, Lawrence. 48pp.
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2016 Taggart, Travis W. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5
2016 Taggart, Travis W. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3
2017 Mardis, Dexter R. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10
2017 Taggart, Travis W. Herp Count: Clark County State Lake. Collinsorum 6(2-3):9
2017 Taggart, Travis W. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8
2018 Houck, Mike Herp Count: Fort Riley Military Installation Collinsorum 7(1):17
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
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8/2/2020 11:08:38 AM