EASTERN COPPERHEAD/BROAD-BANDED COPPERHEAD COMPLEX
Agkistrodon contortrix/laticinctus (Linnaeus, 1766)/(Gloyd & Conant 1934)
ăg-kĭs-trō-dŏn — kŏn-tōr-trĭx/lă-tĭ-sĭnk-tŭs


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None

Adult Eastern Copperhead from Marshall County. © Maci Loughrea.
Adult Eastern Copperhead from Marshall County. © Maci Loughrea.
An adult specimen from Johnson County, Kansas. Image © Suzanne L. Collins, CNAH.
An juvenile from Douglas County. Image © Suzanne L. Collins, CNAH.
An adult Eastern  Copperhead from Cherokee County, Kansas. © Suzanne L. Collins, CNAH.
An adult from Waubaunsee County. Image © Jim Scharosch.
An adult Eastern  Copperhead from Cherokee County, Kansas. © Suzanne L. Collins, CNAH.
An adult from Elk County. Image © Michael Pearce.
An adult Eastern Copperhead basking above Leavenworth State Fishing Lake. © Julie & John Tollefson.
A pair of adults from Wilson County. Image © Adam Sweetman.

Description:
DANGEROUSLY VENOMOUS (a threat to life or limb): Pattern of 7-20 distinct hourglass-shaped light-edged bands with no rattle on the tail and a small pit on each side of its head between and slightly below the eye and nostril. Varies in color from gray to light brown with dark gray to reddish-brown crossbands. Head may be gray, brown, or reddish. The belly is white with large dark gray, brown or black blotches on its edges extending a short distance up onto the sides of the body. Young look like miniature adults, but with yellowish or greenish tails. The two species hybridize across most of Kansas.
Adults normally grow 560-915 mm (22- 36 inches) in TL; largest specimen from Kansas: male (KU 196643) from Jefferson County with TL of 1,020 mm (40 inches) collected by Henry S. Fitch on 13 July 1984; maximum length throughout the range: 53 inches (Conant and Collins, 1998); maximum weight for Kansas specimen is about 400 grams (14 ounces).

Distribution:
The two forms are broadly overlapping. Agkistrodon contortrix is found generally north and east of the Neosho River basin, where it is scarce and occurs in scattered populations. It enters the Flint Hills along rocky riparian corridors where there has been genetic admixture at some point in the past with the Broad-banded Copperhead.
Hybrids occur primarily between the Verdigris River and Walnut River drainages in Kansas.
Additionally, genetic hybrids were identified among the westernmost populations of the Eastern Copperhead (A. contortrix) along the Kansas and Neosho River drainages (Burbrink and Guiher, 2014).
Two 'pure' Eastern Copperheads were detected in Elk County and these outlier samples should be reexamined.

(, Museum Voucher) (, Observation) (, Literature Record)
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  • Occurrence Summary:  
  • 937
    Records 
  • 842
    Museum Vouchers 
  • 95
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (1); Anderson (57); Atchison (18); Bourbon (31); Butler (14); Chase (8); Chautauqua (21); Cherokee (39); Coffey (3); Cowley (28); Crawford (3); Dickinson (6); Doniphan (2); Douglas (164); Elk (14); Franklin (109); Geary (4); Greenwood (7); Jackson (2); Jefferson (30); Johnson (19); Labette (15); Leavenworth (7); Linn (45); Lyon (9); Marshall (21); Miami (127); Montgomery (2); Neosho (1); Osage (18); Pottawatomie (3); Riley (35); Shawnee (3); Unknown (50); Wabaunsee (2); Washington (2); Wilson (4); Woodson (8); Wyandotte (5);

Natural History:
Copperheads prefer open rocky woodland, woodland edge, and meadows adjacent to woodland. The pattern and color of this snake blend well with leaf litter. They are shy and retiring and were it not for their abundance, it would be more difficult to detect. During spring and fall, it is often found on wooded hilltop rock outcrops with a southern exposure. In the summer, they become nocturnal.
Copperheads most commonly breed in April and May following their emergence from brumation. However, copulation has been observed as late as October, and the species is known to store viable sperm for up to a year for delayed fertilization (Fitch, 1960). Gloyd (1934) examined males collected in April, May, June, July, August, and October that all contained active spermatozoa. Fitch (1960) found active spermatozoa in every month snakes were found active (April into November). He further surmised that reproductive activity could occur in any of the months of seasonal activity.
A litter of one to 14 young is born from August to October. Eats insects (especially emerging cicadas), frogs, toads, lizards, small birds, rodents, and other snakes.

Occurrence Activity:
White dates indicate there is at least a single recorded occurrence on that date. The darker blue a date is, the greater the relative number of observations for that date.
Observation Type: (of recorded types)
Remarks:
First reported in Kansas by Hallowell (1856). The earliest existing specimen is from 1879 (MCZ 5844).
Specimens are needed along the western periphery of its range in Kansas to better define the limits of its range there. Additional specimens are also desirable from the Drift Hills in the northeast and throughout the Neosho River drainage basin.
Burt (1935) reported a specimen that was taken 5 miles east of Rock, Cowley County, Kansas, on 16 April 1934.
Gloyd (1969) described Agkistrodon contortrix phaeogaster. United States National Museum no. 165955, adult male, received from Dr. Henry S. Fitch, July 1960; locality 10 miles south of McLouth, Jefferson County, Kansas; collector not stated. He went on to define its distribution as "A. c. phaeogaster occupies the major portion of the Marais des Cygnes-Osage River drainage system in eastern Kansas and central Missouri. It intergrades with A. c. laticinctus in southern Kansas and northern Oklahoma... ".
Burbrink and Guiher (2014) recognized two species A. contortrix (eastern-most populations) and A. laticinctus (western populations) in the United States. The two species mix over eastern Texas and eastern Oklahoma. And, based on relatively extensive sampling in Kansas they discovered that the entire range in Kansas was an admixture of the two species. Because the two species cannot be differentiated in Kansas without examining their genetics, the complex has been grouped into one account.


Bibliography:
1766 Linné, Carl von (=Linneaus). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tomus I Laurentii Salvii, Stockholm, Holmiae, Editio duodecima, reformata (12th Edition). 1-532pp.
1857 Hallowell, Edward. Notice of a collection of reptiles from Kansas and Nebraska presented to the Academy of Natural Sciences, by Doctor Hammond, U. S. A. Proceedings of the Academy of Natural Sciences, Philadelphia 8():238-253
1859 Cope, Edward D. Catalogue of the venomous serpents in the Museum of the Academy of Natural Sciences of Philadelphia, with notes on the families, genera, and species. Proceedings of the Academy of Natural Sciences, Philadelphia 11():332-347
1877 Mozley, Annie E. List of Kansas snakes in the museum of the Kansas State University. Transactions of the Kansas Academy of Science 6():34-35
1882 Yarrow, Henry C. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249
1886 Ebbutt, Percy G. Emigrant Life in Kansas Swan Sonnenschein and Company, Paternoster Square, London. 237pp.
1895 Stejneger, Leonhard. The poisonous snakes of North America. Annual Report of the United States National Museum 1893(2):337-487
1904 Branson, Edwin B. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430
1911 Hurter, Julius. Herpetology of Missouri. Transactions of the Academy of Science St. Louis 20(5):59-274
1913 Crow, H. E. Some trematodes of Kansas snakes. University of Kansas Science Bulletin 7(4):125-136
1925 Linsdale, Jean M. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312pp.
1927 Linsdale, Jean M. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81
1928 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1928 Gloyd, Howard Kay The amphibians and reptiles of Franklin County, Kansas. Transactions of the Kansas Academy of Science 31():115-141
1929 Gloyd, Howard K. Studies on the breeding habits and young of the Copperhead, Agkistrodon mokasen Beauvois, with some observations on ovoviviparity. Thesis. Kansas State Agricultural College, Manhattan, Kansas. 49pp.
1929 Taylor, Edward H. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62
1932 Gloyd, Howard K. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Museum of Zoology, University of Michigan 15():389-408
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1934 Gloyd, Howard K. Studies on the breeding habits and young of the Copperhead, Agkistrodon mokasen Beauvois. Papers of the Michigan Academy of Science, Arts and Letters 19():587-604
1934 Gloyd, Howard K. and Roger Conant. The broad-banded copperhead: A new subspecies of Agkistrodon mokasen. Occasional Papers of the Museum of Zoology, University of Michigan (283):1-5
1935 Burt, Charles E. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16(3):311-366
1935 Burt, Charles E. and William L. Hoyle. Additional records of the reptiles of the central prairie region of the United States Transactions of the Kansas Academy of Science 37():193-216
1938 Gloyd, Howard K. and Roger Conant. The subspecies of the copperhead, Agkistrodon mokasen Beauvois. Bulletin of the Chicago Academy of Sciences 5(7):163-166
1941 Mansueti, R. A descriptive catalogue of the amphibians and reptiles found in and around Baltimore City, Maryland, within a radius of twenty miles. Proceedings of the Natural History Society of Maryland 7():1-53
1942 Hudson, G. E. The amphibians and reptiles of Nebraska. Nebraska Conservation Bulletin 24():1-146
1943 Gloyd, Howard K. and Roger Conant. A synopsis of the American forms of Agkistrodon (copperheads and moccasins). Bulletin of the Chicago Academy of Sciences 7(2):147-170
1946 Gloyd, Howard K. Some rattlesnake dens of South Dakota. The Chicago Naturalist 9(4):87-97
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
1952 Stickel, William H. Venomous snakes of the United States and treatment of their bites. Wildlife Leaflet (339):34
1953 Schmidt, Karl P. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280pp.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
1959 Fitch, Henry S. A patternless phase of the Copperhead. Herpetologica 15(1):21-24
1959 Prophet, Carl W. An outline for conservation teaching in Kansas. Kansas School Naturalist 5(3):16
1960 Fitch, Henry S. Autecology of the copperhead. University of Kansas Publications Museum of Natural History 13(4):85-288
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1969 Gloyd, Howard K. Two additional subspecies of North American crotalid snakes, genus Agkistrodon. Proceedings of the Biological Society of Washington 82():219-232
1970 Fitch, Henry S. Reproductive cycles in lizards and snakes. University of Kansas Museum of Natural History Miscellaneous Publication (52):1-247
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1977 Perry, Janice. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4
1978 Capron, Marty. Notes on the possible occurrence of Broadbanded Copperheads in Kansas. Kansas Herpetological Society Newsletter (26):4-6
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
1978 Fitch, Henry S. and E. Raymond Hall. A 20year record of succession on reseeded fields of tallgrass prairie on the Rockefeller Experimental Tract. University of Kansas Museum of Natural History Special Publication (4):1-15
1978 Perry, Janice. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5
1979 Guarisco, Hank. Preliminary laboratory observatoins of predation by native Texas Garter Snakes upon hatchling Five-lined Skinks. Kansas Herpetological Society Newsletter (32):7-8
1980 Smith, Donald D. The Copperhead. AALAS Kansas City Branch News 3(2):8-10
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
1982 Fitch, Henry S. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C. pp.
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1983 Miller, Larry L. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7
1984 Collins, Joseph T. New records of fishes, amphibians and reptiles in Kansas for 1984. Kansas Herpetological Society Newsletter (58):14-20
1985 Capron, Marty. A Copperhead from Sumner County, Kansas. Kansas Herpetological Society Newsletter (62):11-13
1985 Fitch, Henry S. and Joseph T. Collins. Intergradation of the Osage and Broad-banded Copperheads in Kansas. Transactions of the Kansas Academy of Science 88(3-4):135-137
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1987 Coleman, Keith. Annual KHS Field Trip held at Atchison State Lake. Kansas Herpetological Society Newsletter (68):5-6
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Gloyd, Howard K. and Roger Conant. Snakes of the Agkistrodon complex: A monographic review. Society for the Study of Amphibians and Reptiles Contributions to Herpetology Number 6, . pp.
1990 Lardie, Richard L. Kansas threatened species and protection of the Gypsum Hills habitat. Kansas Herpetological Society Newsletter (80):14-15
1991 Collins, Joseph T Viewpoint: A new taxonomic arrangement for some North American amphibins and reptiles Herpetological Review 22(2):42-43
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Fitch, Henry S. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence. pp.
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Knight, A., L. D. Densmore III, and E. D. Rael. Molecular systematics of the Agkistrodon complex. Pages 49-69 in Biology of the Pitvipers Selva, Tyler, Texas. pp.
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
1994 Fitzgerald, Eve and Charles Nilon Classification of habitats for endangered and threatened species in Wyandotte County, Kansas Kansas Department of Wildlife and Parks, Pratt, Kansas. 98pp.
1994 Riedle, J. Daren. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13
1994 Rundquist, Eric M. 1994 Field Trip Kansas Herpetological Society Newsletter (95):3-4
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1997 Collins, Joseph T. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1998 Collins, Joseph T. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Gamble, Jerre Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1998 Schmidt, Curtis J. Herpetological observations at Cheyenne Bottoms, Barton County, Kansas. Kansas Herpetological Society Newsletter (113):15
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
1999 Taggart, Travis W. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6
2000 Fitch, Henry S. Population structure and biomass of some common snakes in central North America. Scientific Papers of the Natural History Museum University of Kansas (17):1-7
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2000 Taggart, Travis W. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8
2001 Guarisco, Hank. Ode to an ophidion autumn. Kansas Herpetological Society Newsletter (123):19
2001 Kettle, W. Dean, R. H. Hagen, F. deNoyelles, Jr., and E. A. Martinko. The University of Kansas field station and ecological reserves: A half century of research and education. Miscellaneous Publication Number 9. Kansas Biological Survey, Lawrence. 68pp.
2002 Fitch, Henry S. and Anna L. Clark. An exceptionally large natural assemblage of female Copperheads (Agkistrodon contortrix). Herpetological Review 33(2):94-95
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2003 Fitch, Henry S. Reproduction in snakes of the Fitch Natural History Reservation in northeastern Kansas. Journal of Kansas Herpetology (6):21-24
2003 Fitch, Henry S., Scott Sharp, and Kylee Sharp. Snakes of the University of Kansas biotic succession area. Journal of Kansas Herpetology (8):20-21
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Suleiman, Gibran. Fort Riley herp count. Journal of Kansas Herpetology (7):9
2003 Taggart, Travis W. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Taggart, Travis W. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15
2004 Fitch, Henry S. Food surplus and body size in local populations of snakes. Journal of Kansas Herpetology (10):14-16
2004 Fitch, Henry S. Observations on Osage Copperheads in northeastern Kansas. Journal of Kansas Herpetology (12):20
2004 Fitch, Henry S. The effect of female size on number of eggs or young in snakes. Journal of Kansas Herpetology (9):11-12
2004 Volkmann, Al. Cowley County herp count. Journal of Kansas Herpetology (11):10
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2008 Taggart, Travis W. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3
2009 Douglas, Michael E., Marlis R. Douglas, Gordon W. Schuett, and Louis W. Porras. Climate change and evolution of the New World pitviper genus Agkistrodon (Viperidae). Journal of Biogeography 36():1164-1180
2009 Murrow, Daniel G. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Houck, Mike. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9
2011 McMartin, David C. U. S. Army 2011 Fort Leavenworth Herpetofaunal Survey: 23 April - 09 May 2011. Privately printed, Leavenworth, Kansas. 33pp.
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2013 Sinclair, Tom A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9
2013 Taggart, Travis W. KHS 2012 Fall Field Trip to Atchison County State Lake. Collinsorum 2(3/4):4
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6
2013 Taggart, Travis W. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4
2014 Burbrink, Frank T. and Timothy J. Guiher. Considering gene flow when using coalescent methods to delimit lineages of North American pitvipers of the genus Agkistrodon. Zoological Journal of the Linnean Society 2014():1-22
2014 McMartin, D. Chris. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10
2014 Taggart, Travis W. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10
2014 Taggart, Travis W. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12
2015 Mike Houck. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11
2016 Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle Vertebrate animals on the Fitch Natural History Reservation (1948-2002) Kansas Biological Survey Report Number 188, Lawrence. 48pp.
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5
2018 Houck, Mike. Herp Count: Fort Riley Military Installation Collinsorum 7(1):17
2018 Riedle, Daren. Conservation conversations: The year of the snake. Kansas Wildlife and Parks Magazine (November/December):6
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
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9/13/2020 9:57:27 PM