AMPHIBIA (Amphibians) ANURA (Frogs and Toads) RANIDAE (True Frogs)

AMERICAN BULLFROG
Lithobates catesbeianus (Shaw 1802)
lĭth-ō-bā'-tēz — kă-tĕs-bē-ā'-nŭs


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None

An adult female American Bullfrog from Ellis County. Image © Jacob Alexander.
Image © Suzanne L. Collins, CNAH.
A tadpole of Rana catesbieanna. Image © Altig et al. (2006).
A tadpole from Ellis County, Kansas. Image © Travis W. Taggart.
A transforming juvenile from Ellis County, Kansas. Image © Travis W. Taggart.
An adult from an oxbow south of Schermerhorn Park in Cherokee County. Image © Travis W. Taggart.
Common Gartersnake eating an American Bullfrog in Barton County, Kansas. © Ariel Snyder.
A tadpole of Rana catesbieanna. Image © Altig et al. (2006).

Description:
Length in Kansas up to 6.5 inches. This frog is the largest in Kansas. They have a light to dark green ground color with little to heavy dark brown mottling on the head, back, and limbs. They do not have a raised ridge of skin along the sides from the eyes to the legs. The eardrums (tympanum) of males are much larger than the size of the eye.
The largest frog in Kansas. The adults are normally 90-152 mm (3½-6 inches) in snout-vent length. The largest Kansas specimen is a female (KU 181593) from Chase County with a snout-vent length of 185 mm (7¼ inches) collected by Brett Haller on 26 May 1979. The maximum length throughout the range is 8 inches (Conant and Collins, 1998). The heaviest Kansas specimen weighed slightly over 580 grams (about 1 pound, 4 ounces).

Distribution:
Found throughout Kansas but appreciably less common and confined to riparian areas (and nearby impoundments) in the western third of the state. Most likely native to Kansas though probably not found statewide until introduced by fish stocking and as a food source.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record)
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  • Occurrence Summary:  
  • 1,801
    Records 
  • 1,280
    Museum Vouchers 
  • 521
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (15); Anderson (10); Atchison (6); Barber (9); Barton (30); Bourbon (24); Brown (39); Butler (55); Chase (17); Chautauqua (13); Cherokee (86); Cheyenne (8); Clark (9); Clay (20); Cloud (9); Coffey (43); Comanche (9); Cowley (63); Crawford (32); Dickinson (10); Doniphan (17); Douglas (147); Edwards (2); Elk (12); Ellis (20); Ellsworth (21); Finney (8); Ford (5); Franklin (13); Geary (11); Gove (9); Graham (5); Grant (1); Gray (1); Greenwood (47); Hamilton (1); Harper (14); Harvey (37); Hodgeman (3); Jackson (22); Jefferson (18); Jewell (7); Johnson (82); Kingman (17); Kiowa (4); Labette (35); Lane (1); Leavenworth (29); Lincoln (3); Linn (29); Logan (5); Lyon (15); Marion (34); Marshall (5); McPherson (28); Meade (12); Miami (19); Mitchell (1); Montgomery (48); Morris (2); Morton (16); Nemaha (6); Neosho (12); Ness (2); Norton (7); Osage (8); Osborne (1); Ottawa (1); Pawnee (5); Phillips (1); Pottawatomie (32); Pratt (34); Rawlins (3); Reno (15); Republic (4); Rice (7); Riley (35); Rooks (2); Rush (2); Russell (4); Saline (20); Scott (13); Sedgwick (63); Seward (7); Shawnee (38); Sheridan (3); Sherman (5); Smith (1); Stafford (7); Stanton (4); Stevens (1); Sumner (21); Thomas (1); Trego (14); Unknown (18); Wabaunsee (13); Wallace (10); Washington (16); Wichita (1); Wilson (7); Woodson (37); Wyandotte (7);

Natural History:
Frequently seen sitting on vegetation mats at the edge of shallow streams and ponds, or with their eyes just breaking the surface.
They breed on the surface of shallow, permanent water covered with vegetation. Males make distinctive, resonant, low-pitched ‘jug-o-rum’ call both day and night during late spring and through the summer. Females may lay up to 20,000 eggs in one year. The eggs hatch in less than a week and the developing tadpole will remain aquatic for one to three years until transforming into a young terrestrial frog. Bullfrogs are opportunistic predators, and prey on any animal they can fit into their mouths (including smaller American Bullfrogs).
Hartman (1906) removed an adult sparrow from an adult American Bullfrog, and water-beetles and ground-beetles from young specimens.

Occurrence Activity:
The blue dates denote chorusing activity. The red dates are other occurrences. The darker a date is, the greater the relative number of observations for that date.
Chorusing:

Audio recording by Keith Coleman.

Chorusing Phenology: The black outlined dots denote the Julian date (day of the year; 1 January = 1 to 31 December = 365) an observation was made. The thin red line depicts the range of dates between the beginning of the first, and end of the fourth quartile (excluding outliers; Tukey method). The thick light blue bar represents the second and third quartile (interquartile range; the middle 50% of all observations). Only one observation per Julian date is included in the graphs; so a date with multiple observations carries the same weight as a date with only one observation. The vertical bars correspond to the 12 months of the year; January through December.
# Unique Obervations: 396; Range: 11 Mar to 14 Aug; Interquartile range: 17 May to 26 Jun;

Remarks:
First reported in Kansas by Cragin (1880) where he cites correspondence with entomologist Edwin A. Popenoe of Kansas State University "Topeka and Manhattan; rather common; inhabits only deeper streams." The earliest existing specimen (KU 8619) was collected by Frank A. Hartman at Talmage (Dickinson County) on 5 September 1905.
The legs of the American Bullfrog are a delicacy, prized by Kansas sportsmen. They are one of the few herpetological species in Kansas that are regulated as game animals. American Bullfrog season extends from July 1 through October 31 of each year. The daily creel limit is eight. The possession limit is 24 American Bullfrogs after the third day of the season. A valid fishing license is required for any person to take, catch, or kill American Bullfrogs, except persons exempt by law from having such a license. American Bullfrogs may be legally taken any time of day or night by dip net, gig, hook and line, hand, bow, and arrow, or crossbow. A line must attach the bow to an arrow, and the arrow must have a barbed head. Any other method of taking American Bullfrogs is prohibited.

Bibliography:
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1802 Shaw, George General zoology or systematic natural history. Volume 3. G. Kearsley, London, U. K.. 615pp.
Original descriptions of the Rana catesbeiana (=Lithobates catesbeianus) page 106, Lacerta maculata (=Ambystoma maculatum) page 304, and Coluber flagellum (=Masticophis flagellum) page 475.
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():112-123
Also listed the Scarlet Snake (Cemophora coccinea) [=Ophibolus doliatus var. coccineus] and Red Salamander (Pseudotriton ruber) [=Spelerpes ruber] from Kansas.
1886 Ebbutt, Percy G. Emigrant Life in Kansas Swan Sonnenschein and Company, Paternoster Square, London. 237pp.
Life around their homestead, seven miles from Parkerville along the Neosho River, in Morris County, Kansas. A lot of tales so tall as to render doubt on the few plausible scenarios presented. References to the deafening sounds of toads, as well as the abundance of lizards (including chameleons), and snakes. Many references to hunting and killing snakes. Many references to 'rattlesnakes' however, we can't tell what species they are. Includes an illustration of a Texas Horned Lizard and four illustrations of individuals killing snakes.
1906 Hartman, Frank A. Food habits of Kansas lizards and batrachians. Transactions of the Kansas Academy of Science 20():225-229
1914 Dyche, Lewis L. Enemies of fish. Pages 145-158 in Ponds, Pond Fish and Pond Fish Culture State Department Fish and Game Bulletin No. 1, Kansas State Printing Office, Topeka. pp.
1922 O'Roke, Earl C. Frogs and frogging. Transactions of the Kansas Academy of Science 30():448-451
1923 Schumann, Margaretha A. A vestigial maturation cycle in the male larvae of Rana catesbeiana. Thesis. University of Kansas, Lawrence, Kansas. 18pp.
1925 Linsdale, Jean M. Land Vertebrates of a Limited Area in Eastern Kansas. Thesis. University of Kansas, Lawrence, Kansas. 312pp.
1926 Forney, Elsie A. The fauna of an artificial pond. Thesis. University of Kansas, Lawrence. 76pp.
Reported Ambystoma tigrinum, Acris blanchardi, Lithobates blairi, and Lithobates catesbeianus from a man-made well/pond structure just northeast of the junction of East 8th and Pennsylvania Street (ca. 38.969716°, -95.228627°) in northeast Lawrence. Only larvae of the putative Ambystoma tigrinum were mentioned (however larger specimens were said to have contained eggs). It is possible that the larvae were the more common Ambystoma texanum.
1927 Burt, Charles E. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12
Accounts on forty species known from the vicinity of Manhattan, Riley County, Kansas. There have been several scientific names changes since publication, which is understandable... however, some usages cannot be explained by subsequent taxonomic reappraisals(e.g.  Gastrophryne carolinensis for Gastrophryne olivacea). The writer lists Eumeces (=Plestiodon) obsoletus and E. guttulatus yet correctly reasoning that the latter is the young of the former. Within the account of Tantilla gracilis (a common form) the writer mentions that T. nigriceps was reported from Riley County by Branson (1904) but that the specimens at Kansas State were absent at the time of publication. The Prairie Rattlesnake (Crotalus viridis) was first reported from Riley County by Branson (1904) and is reported by the writer based on a specimen in the museum at Kansas State University by Professor E. A. Popenoe. The Prairie Rattlesnake is not currently native to Riley County, and closest reliable localities are 150 to the west.
1927 Linsdale, Jean M. Amphibians and reptiles of Doniphan County, Kansas. Copeia 1927(164):75-81
1932 Gloyd, Howard K. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Museum of Zoology, University of Michigan 15():389-408
First record of Notophthalmus viridescens from Kansas. Second record (after the type locality) of Pseudacris crucifer from Kansas.
1933 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna.
1933 Smith, Hobart M. The Amphibians of Kansas Thesis. University of Kansas, Lawrence. 383pp.
The first full accounting of the twenty-five species of amphibians known to occur in Kansas. Includes Ambystoma maculatum which is currently not included in the Kansas faunal list.
1934 Smith, Hobart M. The Amphibians of Kansas. American Midland Naturalist 15(4):377-527
The formal publication of Hobart Smith's Master's Thesis (Smith 1933), though there are several updated and additions. In addition to the species accounts for all twenty-five species, the paper includes a history of amphibian biology in Kansas and discussions on taxonomy and physiography. 
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1937 Youngstrom, Karl A. Studies on developing behavior in anuran larvae and tadpoles. Dissertation. University of Kansas, Lawrence, Kansas. 118pp.
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1947 Hall, Henry H. and Hobart M. Smith Selected records of reptiles and amphibians from southeastern Kansas Transactions of the Kansas Academy of Science 49(4):447-454
Report on certain Kansas specimens housed in the collection at what is now Pittsburg State University. Included are several species of dubious status today, including Cryptobranchus alleganiensis from the Neosho and Spring rivers (the only specimens from those significant drainages ever documented), Ambystoma maculatum from just north of Pittsburg, Crawford County, Heterodon nasicus from Crawford County, Opheodrys vernalis from Crawford County, Sonora episcopa from Crawford County, Agkistrodon piscivorus from Cherokee County, Crotalus atrox from Crawford County, and Crotalus viridis from Crawford County. They report several significant range extensions including Kinosternon flavescens from Turkey Creek in southeast Cherokee County, Graptemys geographica from just north of Pittsburg, Crotaphytus collaris from near Columbus, Cherokee County, Sceloporus consobrinus from just north of Pittsburg, Phrynosoma cornutum from Cherokee and Crawford counties, Heterodon platirhinos from Cherokee and Crawford counties, Haldea striatula from Crawford County, Sistrurus tergeminus from Crawford County, and a 402 lb Macrochelys temminckii in Cherokee County from just east of Chetopa (Labette County). They allude to the potential for Anaxyrus fowleri to occur in southeast Kansas and for native populations of Crotalus atrox in south central Kansas (in part from the disclosure that John R. Breukelman [then of ESU] had obtained three specimens in Woods County Oklahoma, 3/4 of a mile south of the Kansas line). None of the specimens the paper was based on, exist today.
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1951 Brumwell, Malcolm J. An ecological survey of the Fort Leavenworth Military Reservation American Midland Naturalist 45(1):187-231
Published posthumously. Lieutenant Brumwell died December 14, 1941, as a result of injuries incurred during the Japanese attack on Pearl Harbor. This paper is a condensed version of his thesis for the Master's degree.
1953 Schmidt, Karl P. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280pp.
Schmidt's first edition of his standardized checklist to North American amphibians and reptiles. Includes several specific references to Kansas in the range descriptions.
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1974 Eshelman, Ralph E. Geology and paleontology of the early Pleistocene Belleville Formation of north central Kansas. Dissertation. University of Michigan, Ann Arbor, Michigan. 137pp.
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Perry, Janice. KHS members take trip to southwest Kansas. Kansas Herpetological Society Newsletter (3):2-3
Account of a search for Crotalus atrox and other species discovered in Comanche County.
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1975 Rundquist, Eric M. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3pp.
Short accounts for twenty-nine recognized amphibians and reptiles from Kingman County, Kansas. With habitat descriptions and for some species, estimates of population density.
1975 Rundquist, Eric M. First KHS field trip yields three county records. Kansas Herpetological Society Newsletter (7):1-3
Narration of the activities and species found during the KHS field trip to Kingman County, Kansas. From the title of the article, there were three county records were obtained, however, only Plestiodon septentrionalis is indicated as being 'new'.
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1976 Grow, David. Large garter snake caught at zoo. Kansas Herpetological Society Newsletter (12):8
Winter activity in several herp species on the grounds of the Sedgwick County Zoo.
1976 Grow, David. The KHS goes to Chetopa. Kansas Herpetological Society Newsletter (13):2-3
Spring field trip (22 May 1976) along the Neosho River.
1976 Capron, Marty and Jan Perry. A July weekend in Great Bend. Kansas Herpetological Society Newsletter (14):1-2
1976 Grow, David. A record size Bullfrog. Kansas Herpetological Society Newsletter (16):4
1977 Rundquist, Eric M. and Joseph T. Collins. The amphibians of Cherokee County, Kansas. Kansas Biological Survey, Lawrence. 12pp.
1977 Knight, James L. and Joseph T. Collins. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19pp.
1977 Miller, Larry L. Five days in February. Kansas Herpetological Society Newsletter (18):10-11
1977 Irwin, Kelly J. KHS Ottawa County meeting profitable. Kansas Herpetological Society Newsletter (20):1-2
1977 Perry, Janice. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
Notable mentions: Spotted Salamander, Smooth Green Snake
1978 Warner, M. and R. Wencel. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16
1978 Perry, Janice. KHS successful at Miami County State Lake. Kansas Herpetological Society Newsletter (27):5
1979 Miller, Larry L. Life in Kansas and Oklahoma once again threatened by pesticides. Kansas Herpetological Society Newsletter (31):7
Report on damage to wildlife as a result of the aerial spraying of toxaphene and parthion in south-central Kansas for Army-cutworms. Animals noted to be affected (besides cutworms) "...amphibians such as leopard frogs, bullfrogs, and several species of toads. Several turtles, snakes, and other reptiles were also found dead in the area as well as birds and small mammals...".
1979 Guarisco, Hank. Preliminary laboratory observatoins of predation by native Texas Garter Snakes upon hatchling Five-lined Skinks. Kansas Herpetological Society Newsletter (32):7-8
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilisThamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1982 Collins, Joseph T. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57pp.
1982 Fitch, Henry S. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C. pp.
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1983 Miller, Larry L. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26
Invalidated the specimens of Thamnophis sirtalis from Hamilton County (reidentified as T. cyrtopsis; KU 2088) and Wallace County mapped in Collins, 1982.
1985 Guarisco, Hank Opportunistic scavenging by the Bullfrog, Rana catesbeiana (Amphibia, Anura, Ranidae). Transactions of the Kansas Academy of Science 88():38-39
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4
1986 Capron, Marty. If its December it must be time for frogs. Kansas Herpetological Society Newsletter (66):17
1987 Coleman, Keith. Annual KHS Field Trip held at Atchison State Lake. Kansas Herpetological Society Newsletter (68):5-6
1987 Simmons, John E. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894
1988 Schwalbe, Cecil R. and Philip C. Rosen. Preliminary report on effect of Bullfrogs on wetland herpetofaunas in southeastern Arizona. Pages 166-173 in Management of Amphibians, Reptiles, and Small Mammals in North America USDA Forest Service, General Technical Report RM-166. pp.
1988 Dalrymple, George H. The herpetofauna of Long Pine Key, Everglades National Park, in relation to vegetation and hydrology. Pages 72-86 in Management of Amphibians, Reptiles, and Small Mammals in North America. USDA Forest Service, General Technical Report RM-166. pp.
1988 Miller, Larry L. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1988. Kansas Herpetological Society Newsletter (75):15-18
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Collins, Joseph T. and Suzanne L. Collins. Reptiles and Amphibians of the Cimarron National Grasslands, Morton County, Kansas. U. S. Forest Service, Elkhart, Kansas. 60pp.
1991 Fitch, Henry S. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence. pp.
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1992 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Taggart, Travis W. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1994 Riedle, J. Daren. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13
1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4
See, Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.
1995 Anderson, Lewis, Mark Shaw, Jeff Blodig, and Tom Walker. Report to the Kansas Department of Wildlife and Parks: Herps encountered during REmap project, summer 1994. Kansas Herpetological Society Newsletter (99):10-17
1995 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1994. Kansas Herpetological Society Newsletter (100):24-47
1995 Moriarty, Emily C. and Joseph T. Collins. First known occurrence of amphibian species in Kansas. Kansas Herpetological Society Newsletter (100):28-30
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1996 Miller, Larry L. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Miller, Larry L. Third graders conduct amphibian and reptile field study. Kansas Herpetological Society Newsletter (106):15
1996 Miller, Larry L. Many amphibian and reptile species identified during KHS 1996 fall field trip to Wabaunsee County. Kansas Herpetological Society Newsletter (106):2-3
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1997 Rundquist, Eric M. Addendum to 1997 KHS herp counts. Kansas Herpetological Society Newsletter (109):14-15
1997 Taggart, Travis W. Status of Bufo debilis (Anura: Bufonidae) in Kansas Kansas Herpetological Society Newsletter (109):7-12
The specimens (KU) cited from Hamilton County, have been corrected.
1997 Collins, Joseph T. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Gamble, Jerre Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1998 Collins, Joseph T. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
2000 Taggart, Travis W. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5
2000 Van Doren, Mark D. and Curtis J. Schmidt. A herpetological survey of the Fort Larned National Historic Site, Pawnee County, Kansas. Kansas Herpetological Society Newsletter (120):8-11
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2000 Taggart, Travis W. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8
2001 Fitch, Henry S. Further study of the garter snake, Thamnophis sirtalis, in northeastern Kansas. Scientific Papers of the Natural History Museum University of Kansas (19):1-6
2001 Taggart, Travis W. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14
2001 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 2000. Kansas Herpetological Society Newsletter (124):6-8
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Palenske, Nicole M. Blood Viscosity and Hematological Parameters in Hibernating Bullfrogs, Rana catesbeiana. Thesis. Emporia State University, Emporia, Kansas. 61pp.
2002 Riedle, J. Daren and A. Hynek. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20
2002 Ellis, Mark R. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5
2002 Taggart, Travis W. Results of the spring 2002 KHS field trip. Journal of Kansas Herpetology (3):6-7
To the Cimarron National Grassland, in Morton and Stevens counties.
2002 Taggart, Travis W. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13
To Washington County, Kansas (also includes ancillary counts from Clay and Marshall counties).
2002 Miller, Larry L. Osage County herp count II. Journal of Kansas Herpetology (4):15
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Taggart, Travis W. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (7):10
2003 Miller, Larry L. and Suzanne L. Miller. Wakarusa herp count. Journal of Kansas Herpetology (7):10
2003 Burr, Andrew. Coffey County herp count 1. Journal of Kansas Herpetology (7):7
2003 Volkmann, Al. Cowley County herp count 1. Journal of Kansas Herpetology (7):7
2003 Gubanyi, James E. Osage County herp count. Journal of Kansas Herpetology (7):8
2003 Lokke, John L. and Jill Lokke. Cowley County herp count 2. Journal of Kansas Herpetology (7):8
2003 Gubanyi, James E. Shawnee County herp count. Journal of Kansas Herpetology (7):9
2003 Miller, Larry L. Indian Creek herp count. Journal of Kansas Herpetology (7):9
2003 Suleiman, Gibran. Fort Riley herp count. Journal of Kansas Herpetology (7):9
2003 Taggart, Travis W. Results of the KHS 2003 fall field trip. Journal of Kansas Herpetology (8):14-15
2004 Delisle, Jennifer M. and William H. Busby Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (11):11-12
2005 Taggart, Travis W., Curtis J. Schmidt, and Richard S. Hayes. Geographic distribution: Rana catesbeiana. Journal of Kansas Herpetology (13):10
2005 Flowers, T. L. Life history notes: Rana catesbeiana. Diet. Journal of Kansas Herpetology (14):10
2005 Taggart, Travis W. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21
2005 Hillis, David M. and Thomas P. Wilcox Phylogeny of the New World True Frogs (Rana) Molecular Phylogenetics and Evolution 34(2):299-314
2006 Altig, Ronald, Roy W. McDiarmid, Kimberly A. Nichols, and Paul C. Ustach Tadpoles of the United States and Canada: A Tutorial and Key Electronic files accessible at http://www.pwrc.usgs.gov/tadpole/. Patuxent Wildlife Research Center, Laurel, MD, USA. ():
2006 Taggart, Travis W. Addendum report to biological inventory of the sandsage prairie near Holcomb, Kansas. Sunflower Electric Cooperative, Hays, Kansas. 31pp.
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Frost, Darrel R, Taran Grant, Julian Faivovich, Raoul H. Bain, Alexander Haas, Celio F. B. Haddad, Rafael O. De Sa, Alan Channing, Mark Wilkinson, Stephen C. Donnellan, Christopher J. Raxworthy, Jonathan A. Campbell, Boris L. Blotto, Paul Moler, Robert C. Drewes, Ronald A. Nussbaum, John D. Lynch, David M. Green, and Ward C. Wheeler. The amphibian tree of life Bulletin of the American Museum of Natural History (297):370
2006 Anderson, Lewis R. and Joseph A. Arruda. Land use and anuran biodiversity in southeast Kansas, USA. Amphibian and Reptile Conservation 4(1):46-59
2007 Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5pp.
2007 Schmidt, Curtis J. First record of the Bullfrog from Osborne County, Kansas. Journal of Kansas Herpetology (22):11
2007 Taggart, Travis W. A biological inventory of the Sunflower Electric Site near Holcomb, Kansas. Journal of Kansas Herpetology 23():11-16
2008 Schwenke, Zachery J. Geographic distribution: Lithobates catesbeianus. Kansas. Journal of Kansas Herpetology (27):7
2009 Murrow, Daniel G. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769
2010 Miller, Larry L. Investigation of the Checkered Garter Snake in Kansas with notes on other Amphibians, Reptiles, and Turtles encountered. Kansas Department of Wildlife and Parks, Pratt, Kansas. 31pp.
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 McMartin, David C. U. S. Army 2011 Fort Leavenworth Herpetofaunal Survey: 23 April - 09 May 2011. Privately printed, Leavenworth, Kansas. 33pp.
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Fall Field Trip to Lovewell State Park Journal of Kansas Herpetology (39):4-5
2011 McMartin, D. Chris. Herp Count: Fort Leavenworth Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):8-9
2011 Houck, Mike. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2013 Bass, Neil The Missouri River Fish and Wildlife Mitigation Project: For the river, for you, and for herps. Collinsorum 2(1/2):10-11
2013 Miller, Larry L. Wellington Lake Herpetological Survey. Collinsorum 2(1/2):12
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2012 Summer Field Trip to Meade County State Park. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2012 Fall Field Trip to Atchison County State Lake. Collinsorum 2(3/4):4
2013 Taggart, Travis W. KHS 2013 Summer Field Trip to Coldwater Lake, Comanche County. Collinsorum 2(3/4):5
2013 Taggart, Travis W. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6
2014 McMartin, D. Chris. Fort Leavenworth Heretofaunal Survey for 2013. Collinsorum 3(1):10
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2014 Taggart, Travis W. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12
2014 Taggart, Travis W. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10
2015 Mike Houck. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11
2015 Bass, Neil. Herpetological (Frog and Turtle) Inventories along the Missouri River in Kansas. Collinsorum 4(1):5-9
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2015 Taggart, Travis W. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3
2015 Taggart, Travis W. Fall Field Trip Held In Washington County. Collinsorum 4(3):4
2016 Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle Vertebrate animals on the Fitch Natural History Reservation (1948-2002) Kansas Biological Survey Report Number 188, Lawrence. 48pp.
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3
2016 Taggart, Travis W. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2017 Snyder, Ariel. Survey Of Anuran Chytrid (Batrachochytrium dendrobatidis) in Kansas and the Influence of Anuran Life History in Occurrence. Thesis. Fort Hays State University, Hays, Kansas. 53pp.
2017 Taggart, Travis W. and J. Daren Riedle. A Pocket Guide to Kansas Amphibians, Turtles and Lizards. Great Plains Nature Center, Wichita, Kansas. 69pp.
2017 Mardis, Dexter R. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10
2017 Taggart, Travis W. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8
2018 Houck, Mike. Herp Count: Fort Riley Military Installation Collinsorum 7(1):17
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Neosho County: KHS-2020-01 Collinsorum 9(3):11
2021 Mardis, Dexter R. Herp Count: Sumner County: KHS-2020-08. Collinsorum 9(3):12-13
2021 Taggart, Meg, Amelia Jaeger, Jesse J. Taggart, and Travis W. Taggart. Herp Count: Russell County: KHS-2020-22. Collinsorum 9(3):14-15
2021 Taggart, Megan M. and Travis W. Taggart. Herp Count: Seward County: KHS-2020-27. Collinsorum 9(3):15
2021 Riedle, J. Daren. Herp Count: Pratt County: KHS-2020-33. Collinsorum 9(3):16
2021 Dodd, C. Kenneth, Jr. and Mark R. Jennings. How to raise a Bullfrog - The literature on frog farming in North America. Bibliotheca Herpetologica 15(8):1-24
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5/26/2021 7:11:22 AM