An adult female Plains Spadefoot from Wallace County, Kansas. © Travis W. Taggart.
An adult male Plains Spadefoot chorusing in Pratt County, Kansas. © Travis W. Taggart.
An adult Plains Spadefoot from Scott County. © Kurt Meier.
An adult female Plains Spadefoot from Morton County, Kansas. © Travis W. Taggart.
An adult female Plains Spadefoot from Logan County, Kansas. © Travis W. Taggart.
A tadpole of Spea bombifrons. Image © Altig et al. (2006).
AMPHIBIA (Amphibians) ANURA (Frogs) SCAPHIOPODIDAE (North American Spadefoots)

Plains Spadefoot
Spea bombifrons (Cope 1863)
spē-ŭh — bäm-bĭ'-fräns


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
The Plains Spadefoot is characterized by moist skin, a round snout, eyes which are vertically slit when exposed to strong light, and a black spur at the base of each hind foot. The head, body, and limbs of this species are light to medium gray with irregular darker markings and two poorly defined light lines down the back. The belly is white. Males have dark throats, and females have heavier bodies than males.
Adults normally 38-50 mm (1½-2 inches) in snout-vent length. The largest Kansas specimen is a male (KU 20012) from Barber County with snout-vent length of 64 mm (2½ inches) collected by Claude W. Hibbard on 29 August 1935. This is the maximum length throughout the range (Powell et al., 2016).

Distribution:
The Plains Spadefoot is found in western Kansas to the edge of the Flint Hills, and east along the floodplain of the Kansas and Missouri rivers. It is seldom encountered except when chorusing or moving (often across roads) The Plains Spadefoot inhabits prairies and open floodplains. It prefers areas of loose soil or sand, into which during rainy weather it burrows during daylight hours and from which it emerges in the evening to forage, particularly after rain. In northeastern Kansas, Fitch (1958) found this species scarce in the wooded uplands of his study area and considered small mammal burrows to be their favored retreats. Plains Spadefoots are active from late April to September.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
Open icons are questionable records; Click on a marker to view details.
Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 1,976
    Records 
  • 1,744
    Museum Vouchers 
  • 232
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Atchison (3); Barber (86); Barton (33); Cheyenne (19); Clark (16); Clay (2); Cloud (2); Comanche (32); Cowley (7); Decatur (6); Dickinson (1); Doniphan (1); Douglas (221); Edwards (3); Ellis (130); Ellsworth (1); Finney (86); Ford (16); Geary (2); Gove (1); Graham (18); Grant (7); Gray (11); Greeley (5); Hamilton (9); Harper (36); Harvey (3); Haskell (7); Hodgeman (1); Jefferson (32); Jewell (3); Johnson (12); Kearney (12); Kingman (38); Kiowa (2); Lane (6); Leavenworth (2); Logan (40); Marion (1); Marshall (5); McPherson (19); Meade (82); Mitchell (7); Morton (159); Ness (27); Norton (1); Osborne (5); Ottawa (1); Pawnee (20); Phillips (10); Pottawatomie (15); Pratt (34); Rawlins (1); Reno (70); Rice (9); Riley (39); Rooks (7); Rush (161); Russell (10); Saline (4); Scott (30); Sedgwick (4); Seward (27); Shawnee (2); Sheridan (2); Sherman (53); Smith (2); Stafford (72); Stanton (2); Stevens (7); Sumner (5); Thomas (119); Trego (14); Unknown (2); Wabaunsee (3); Wallace (24); Washington (3); Wichita (2); Wyandotte (4);

Fossil History:
Pleistocene fossils of this species have been collected from McPherson and Meade counties.
Fossils from the Sandahl Local Fauna of McPherson County (Pleistocene: Illinoian) (Holman 1971; Preston 1979) are assignable to this taxon.

Natural History:
The Plains Spadefoot is an explosive and opportunistic breeder, emerging from its underground retreats for this purpose after the first warm, heavy spring rain and continuing to congregate under suitable conditions throughout the summer. Males loudly chorus for females to join them around temporary rain-pools on floodplains or open prairie. The male mounts a female, clasping her around the groin with his front limbs. The female deposits her eggs, and the male arches his body and fertilizes them in the water. Each female may lay up to 2,000 eggs in masses of 10- 250 each; they are attached to partly submerged vegetation or other protruding objects.
The eggs hatch, and metamorphosis occurs at widely varying intervals dependent on water temperature, oxygen content, and competition for available food between tadpoles. Evidently, tadpoles of this species are cannibalistic under crowded conditions. Smith (1934) stated that this species breeds anytime after the first heavy rain of mid-spring and (1956) specified that this species normally breeds after a minimum spring rainfall of 88.9 mm (3.5 inches) coincident with an air temperature of 52°F or higher. Fitch (1958) reported that this species breeds chiefly on the lowland floodplain of the Kansas River in Douglas County. Hoyt (1960) found adults mating in a temporary pool on 2 July in Riley County and gave a detailed description of egg-laying and eggs of this species. Knight and Collins (1977) reported adult males of the Plains Spadefoot calling on 23 May at 2300 hours amidst a chorus of Woodhouse's Toads in a sandpit lake in Cheyenne County and observed large numbers of tadpoles of the Plains Spadefoot during daylight hours in small impoundments in the same county. Heinrich and Kaufman (1985) reported two males calling from a temporary pool in a roadside ditch on 10 June in the Konza Prairie. Irwin and Collins (1987) reported two of these amphibians active at night on 4 May and heard seven Plains Spadefoots chorusing at night on 2 June, all at Cheyenne Bottoms in Barton County.
Beetles, crickets, grasshoppers, ants, and other small insects are the preferred food of this species.
Brennan (1936) reported predation on a young Plains Spadefoot by a domestic cat in Ellis County.

Occurrence Activity:
Number of Unique Obervations (=days): 159; Range: 02 Jan to 16 Oct
Chorusing:

Audio recording by Keith Coleman.

Chorusing Phenology: The black outlined dots denote the Julian date (day of the year; 1 January = 1 to 31 December = 365) an observation was made. The thin red line depicts the range of dates between the beginning of the first, and end of the fourth quartile (excluding outliers; Tukey method). The thick light blue bar represents the second and third quartile (interquartile range; the middle 50% of all observations). Only one observation per Julian date is included in the graphs; so a date with multiple observations carries the same weight as a date with only one observation. The vertical bars correspond to the 12 months of the year; January through December.
Number of Unique Obervations: 165; Range: 27 Mar to 14 Aug; Interquartile range: 10 May to 27 Jun;

Remarks:
First reported from Kansas by Taylor (1929) based on more than 50 specimens taken on both 18 August 1926 after heavy rainfall in Morton County. The oldest available specimen from Kansas is KU 5211, collected 18 miles north of Elkhart, Morton County, on 10 August 1926 by Theodore E. White and Edward H. Taylor.
The discrepancy between Taylor's publication (1929) and the respective catalogs of Morton County specimens collected in 1926 are noted. There are 57 Spea bombifrons from "18 mi N Elkhart" collected on 10 August 1926 (KU 5211-67) by Theodore E. White and Edward H. Taylor. There are three specimens (BYU 2159, 2739; FMNH 103461) collected " mi. N Elkhart" on 15 August 1926 by Theodore E. White and Edward H. Taylor. Four specimens (KU 7179-82) collected "41 Ranch, Rolla" on 19 August 1926. KU 3523 from "Elkhart, 81 Ranch" on 29 June 1926.KU 5208 from "18 mi N Elkhart" on 29 June 1926.
Plains Spadefoots are active on humid nights, and during such times, they are often readily encountered on roads through suitable habitat, from April into September.

Bibliography:
1929 Taylor, Edward H. List of reptiles and batrachians of Morton County, Kansas, reporting species new to the state fauna. University of Kansas Science Bulletin 19(6):63-65
Annotated listing of 25 species discovered in southwest Morton County just prior to the Dust Bowl that wreaked havoc on the region and the subsequent creation of the Cimarron National Grassland. Of special note are Heterodon platirhinos, Thamnophis marcianus, and Anaxyrus debilis (all of which have not been found in the area since).
1932 Kellogg, Remington. Notes on the spadefoot of the Western Plains (Scaphiopus hammondii). Copeia 1932(1):36
1933 Smith, Hobart M. The Amphibians of Kansas. Thesis. University of Kansas, Lawrence. 383pp.
The first full accounting of the twenty-five species of amphibians known to occur in Kansas. Includes Ambystoma maculatum which is currently not included in the Kansas faunal list.
1934 Smith, Hobart M. The Amphibians of Kansas. American Midland Naturalist 15(4):377-527
The formal publication of Hobart Smith's Master's Thesis (Smith 1933), though there are several updated and additions. In addition to the species accounts for all twenty-five species, the paper includes a history of amphibian biology in Kansas and discussions on taxonomy and physiography. 
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1935 Brennan, Lawrence A. Notes on the distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1937 Trowbridge, Albert H. and Minnie S. Trowbridge. Notes on the cleavage rate of Scaphiopus bombifrons Cope, with additional remarks on certain aspects of its life history. The American Naturalist 71(736):460-480
1939 Trowbridge, Minnie S. Embryonic and larval development of Scaphiopus bombifrons Cope. Dissertation. University of Oklahoma, Norman. 159pp.
1939 Tihen, Joseph A. and James M. Sprague. Amphibians, reptiles, and mammals of the Meade County State Park Transactions of the Kansas Academy of Science 42():499-512
1941 Trowbridge, Minnie S. Studies on the normal development of Scaphiopus bombifrons. I. The cleavage period. Transactions of the American Microscopical Society 60(4):508-526
1942 Hudson, G. E. The amphibians and reptiles of Nebraska. Nebraska Conservation Bulletin 24():1-146
1942 Trowbridge, Minnie S. Studies on the normal development of Scaphiopus bombifrons. II. The later embryonic and larval periods. Transactions of the American Microscopical Society 61(1):66-83
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1945 Lane, Henry H. A survey of the fossil vertebrates of Kansas, Part II. Amphibia. Transactions of the Kansas Academy of Science 48(3):286-316
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1954 Tihen, Joseph A. A Kansas Pleistocene herpetofauna. Copeia 1954(3):217-221
Kingsdown Formation in Meade County.
1955 Hibbard, Claude W. The Jinglebob Interglacial (Sangamon?) Fauna from Kansas and its climatic significance. Contributions from the Museum of Paleontology, The University of Michigan 12(10):179-228
1956 Chrapliwy, Pete S. Taxonomy and distribution of the spadefoot toads of North America (Salientia: Pelobatidae). Thesis. University of Kansas, Lawrence. 131pp.
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 speciesfor chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1956 Zweifel, Richard G. Two Pelobatid frogs from the Tertiary of North America and their relationship to fossil and recent forms. American Museum Novitates 1762():1-45
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1965 Smith, Hobart M., T. Paul Maslin, and Robert L. Brown. Summary of the distribution of the herpetofauna of Colorado: A Supplement to An Annotated Check List of the Amphibians and Reptiles of Colorado. University of Colorado Studies Series in Biology (15):56
1966 Kluge, Arnold G. A new pelobatine frog from the lower Miocene of South Dakota with a discussion of the evolution of the Scaphiopus/Spea complex. Los Angeles County Museum Contributions in Science 113():42761
1967 Choate, Jerry R. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46pp.
1967 Gier, Herschel T. Vertebrates of the Flint Hills. Transactions of the Kansas Academy of Science 70(1):51-59
1971 Holman, J. Alan. Herpetofauna of the sandahl local fauna (Pleistocene: Illinoian) of Kansas. Contributions from the Museum of Paleontology, University of Michigan 23(22):349-355
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Perry, Janice. KHS members take trip to southwest Kansas. Kansas Herpetological Society Newsletter (3):2-3
Account of a search for Crotalus atrox and other species discovered in Comanche County.
1975 Rundquist, Eric M. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3pp.
Short accounts for twenty-nine recognized amphibians and reptiles from Kingman County, Kansas. With habitat descriptions and for some species, estimates of population density.
1975 Rundquist, Eric M. First KHS field trip yields three county records. Kansas Herpetological Society Newsletter (7):1-3
Narration of the activities and species found during the KHS field trip to Kingman County, Kansas. From the title of the article, there were three county records were obtained, however, only Plestiodon septentrionalis is indicated as being 'new'.
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Capron, Marty B. and Jan Perry. A July weekend in Great Bend. Kansas Herpetological Society Newsletter (14):1-2
1976 Brown, Herbert A. The status of California and Arizona populations of the western spadefoot toads (genus Scaphiopus). Natural History Museum of Los Angeles County, Contributions in Science (286):1-14
Includes a range map of Spea bombifrons and Spea hammondii (Spea multiplicata now) in, or near, Kansas.
1977 Knight, James L. and Joseph T. Collins. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19pp.
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
Notable mentions: Spotted Salamander, Smooth Green Snake
1978 Warner, M. and R. Wencel. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16
1978 Capron, Marty B. Four county collecting raid: A south central Kansas herping saga. Kansas Herpetological Society Newsletter (26):9-12
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1979 Martin, Larry D. Survey of fossil vertebrates from east-central Kansas: Kansas River bank stabilization study. U.S. Army Corps of Engineers, Kansas City District. 55pp.
1979 Ports, Mark A. Occurrence and density studies of nongame wildlife in southwestern Kansas - May 16-August 16, 1979. Kansas Fish and Game Commission, Pratt, Kansas. 83pp.
1979 Gray, Peter and Eddie Stegall. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8
1979 Holman, J. Alan. Herpetofauna of the Nash local fauna (Pleistocene: Aftonian) of Kansas. Copeia 1979(4):747-749
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilisThamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1980 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survery of Kansas 9():1-11
1981 Eshelman, Ralph E. and Claude W. Hibbard. Nash Local Fauna (Pleistocene: Aftonian) of Meade County, Kansas. Contributions from the Museum of Paleontology, The University of Michigan 25(16):317-326
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1983 Trott, Gene. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (52):3-4
1984 Heinrich, Mark L. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57pp.
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1984 Schwarting, Nancy. KHS field trip, May 1984. Kansas Herpetological Society Newsletter (57):3-4
1985 Lynch, John D. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Science 13():33-57
1985 Heinrich, Mark L. and D. W. Kaufman. Herpetofauna of the Konza Prairie Research Natural Area, Kansas Prairie Naturalist 17(2):101-112
1985 Sattler, P. W. Introgressive hybridization between the spadefoot toads Scaphiopus bombifrons and S. multiplicatus (Salientia: Pelobatidae). Copeia 1985():324-332
1987 Irwin, Kelly J. and Joseph T. Collins. Amphibians and Reptiles of Cheyenne Bottoms. Pages 401-432 in Cheyenne Bottoms: An Environmental Assessment. Kansas Biological Survey and Kansas Geological Survey, Pratt, Kansas. pp.
1988 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19
1989 Wiens, John J. Ontogeny of the skeleton of Spea bombifrons (Anura: Pelobatidae). Journal of Morphology 202():29-51
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Collins, Joseph T. and Suzanne L. Collins. Reptiles and Amphibians of the Cimarron National Grasslands, Morton County, Kansas. U. S. Forest Service, Elkhart, Kansas. 60pp.
1991 Fitch, Henry S. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas. pp.
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1991 Wiens, John J. and Tom A. Titus. A phylogenetic analysis of Spea (Anura: Pelobatidae). Herpetologica 47(1):21-28
1992 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Taggart, Travis W. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4
1992 Taggart, Travis W. KHS field trips. Kansas Herpetological Society Newsletter (91):3
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1994 Fitzgerald, Eve C. and Charles Nilon. Classification of habitats for endangered and threatened species in Wyandotte County, Kansas Kansas Department of Wildlife and Parks, Pratt, Kansas. 98pp.
1994 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19
1995 Holman, J. Alan. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243pp.
1995 Moriarty, Emily C. and Joseph T. Collins. First known occurrence of amphibian species in Kansas. Kansas Herpetological Society Newsletter (100):28-30
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1996 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1995. Kansas Herpetological Society Newsletter (103):13-15
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1997 Taggart, Travis W. Status of Bufo debilis (Anura: Bufonidae) in Kansas Kansas Herpetological Society Newsletter (109):7-12
The specimens (KU) cited from Hamilton County, have been corrected.
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
2000 Maglia, Anne M. Phylogenetic Relationships of Pelobatoid Frogs (Anura: Pelobatoidea). Dissertation. University of Kansas, Lawrence, Kansas. pp.
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2001 Taggart, Travis W. Results of the KHS spring field trip west. Kansas Herpetological Society Newsletter (125):10
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2001 Kretzer, Justin E. and Jack F. Cully, Jr. Effects of Blacktailed Prairie Dogs on reptiles and amphibians in Kansas shortgrass prairie. Southwestern Naturalist 46(2):171-177
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Fogell, Daniel D. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6pp.
2002 Ellis, Mark R. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Taggart, Travis W. Logan County herp count. Journal of Kansas Herpetology (7):8
2004 Schmidt, Curtis J. Natural history and status of the exploited Prairie Rattlesnake (Crotalus viridis) in western Kansas and a herpetofaunal inventory of the Smoky Valley Ranch, Logan County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. pp.
2004 Delisle, Jennifer M. and William H. Busby. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Taggart, Travis W. Kansas Herpetological Society 2004 spring field trip. Journal of Kansas Herpetology (9):2
2005 Lannoo, Michael (Editor) Amphibian Declines: The Conservation Status of United States Species. University fo California Press, Berkeley. 1115pp.
2006 Altig, Ronald, Roy W. McDiarmid, Kimberly A. Nichols, and Paul C. Ustach. Tadpoles of the United States and Canada: A Tutorial and Key Electronic files accessible at http://www.pwrc.usgs.gov/tadpole/. Patuxent Wildlife Research Center, Laurel, MD, USA. ():
2006 Taggart, Travis W. Addendum report to biological inventory of the sandsage prairie near Holcomb, Kansas. Sunflower Electric Cooperative, Hays, Kansas. 31pp.
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Frost, Darrel R, Taran Grant, Julian Faivovich, Raoul H. Bain, Alexander Haas, Celio F. B. Haddad, Rafael O. De Sa, Alan Channing, Mark Wilkinson, Stephen C. Donnellan, Christopher J. Raxworthy, Jonathan A. Campbell, Boris L. Blotto, Paul Moler, Robert C. Drewes, Ronald A. Nussbaum, John D. Lynch, David M. Green, and Ward C. Wheeler. The amphibian tree of life Bulletin of the American Museum of Natural History (297):370
2007 Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5pp.
2007 Taggart, Travis W. A biological inventory of the Sunflower Electric Site near Holcomb, Kansas. Journal of Kansas Herpetology 23():11-16
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2011 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of the Cimarron National Grassland, Kansas. Second (Revised) Edition. U. S. Forest Service, . pp.
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. and Daniel Murrow. KHS to conduct summer field trip to western Kansas. Journal of Kansas Herpetology (38):5
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2013 Dodd, C. Kenneth. Frogs of the United States and Canada. John Hopkins University Press, Baltimore, Maryland. 982pp.
2013 Bass, Neil. The Missouri River Fish and Wildlife Mitigation Project: For the river, for you, and for herps. Collinsorum 2(1/2):10-11
2013 Taggart, Travis W. KHS 2012 Summer Field Trip to Meade County State Park. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Summer Field Trip to Coldwater Lake, Comanche County. Collinsorum 2(3/4):5
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2014 Taggart, Travis W. Results of the 2014 KHS summer field trip to Morton County and adjacent Colorado, New Mexico, and Oklahoma. Collinsorum 3(2-4):12
2014 Pence, William S. Reconfirmation of the Plains Spadefoot (Spea bombifrons) at the Fort Riley Military Reservation. Collinsorum 3(2-4):13
2015 Brown, Kasandra A. Occupancy Modeling Of Herpetofauna And Grassland Nesting Birds At Quivira National Wildlife Refuge. Thesis. Fort Hays State University, Hays, Kansas. 72pp.
2015 Bass, Neil. Herpetological (Frog and Turtle) Inventories along the Missouri River in Kansas. Collinsorum 4(1):5-9
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2015 Taggart, Travis W. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3
2016 Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle. Vertebrate animals on the Fitch Natural History Reservation (1948-2002) Kansas Biological Survey Report Number 188, Lawrence. 48pp.
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2017 Snyder, Ariel. Survey Of Anuran Chytrid (Batrachochytrium dendrobatidis) in Kansas and the Influence of Anuran Life History in Occurrence. Thesis. Fort Hays State University, Hays, Kansas. 53pp.
2017 Taggart, Travis W. and J. Daren Riedle. A Pocket Guide to Kansas Amphibians, Turtles and Lizards. Great Plains Nature Center, Wichita, Kansas. 69pp.
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2021 Taggart, Travis W. Herp Count: Harper County: KHS-2020-26. Collinsorum 9(3):15
2023 Dodd, C. Kenneth. Frogs of the United States and Canada. Second Edition. John Hopkins University Press, Baltimore, Maryland. 1032pp.
2023 Russell, Elisabeth Habitat associations and fine-scale movements of the Red-spotted Toad (Anaxyrus punctatus) in Kansas and the efficacy of remote telemetry for monitoring small-scale movements. Thesis. Fort Hays State University, Hays, Kansas. 81pp.
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Travis W. Taggart © 1999-2025 — w/ Sternberg Museum of Natural History, Fort Hays State University