Kansas Herpetofaunal Atlas

An adult Lined Snake from Montgomery County, Kansas. Image © Daren Riedle.
The belly of an adult Lined Snake from Elk County. Image © Mary Liz Jameson (iNat).
An adult from Waubaunsee County. Image © Jim Scharosch.
An adult Lined Snake from Ottawa County. Image © Anna Dawson (iNat).
An adult Lined Snake from Wabaunsee County. Image © Michael Tobler (iNat).
An adult Lined Snake from Riley County. Image © Michael Tobler (iNat).
An adult Lined Snake from Sumner County. Image © Sylbird (iNat).
An adult from Morris County. Image © Suzanne L. Collins, CNAH.
REPTILIA (Reptiles) SQUAMATA (PART) (Snakes) NATRICIDAE (Harmless Egg-Retaining Snakes)

Lined Snake
Tropidoclonion lineatum (Hallowell 1856)
trŏp-ĕh-dō-klōn-ē-ŏn — lĭn-ē-ā-tŭm

Conservation Status:
State: None
Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
 HARMLESS. The Lined Snake is characterized by keeled scales, a single anal scale, six or fewer scales on each upper lip, and a white or yellowish belly with a double row of black spots down the middle. The head, body, and tail are gray or brown. There are three gray-white to yellowish stripes (one down the middle of the back and one on each side) on the body. Adult males have longer tails than females.
The body scales are keeled and the anal scale is single. There are six or fewer scales on each upper lip. The young look like tiny adults.
Adults normally grow 22.4-38.0 cm (8­5⁄8-15 inches) in total length. The largest specimen from Kansas is a female (KU 208123) from Sedgwick County with a total length of 44.6 cm (17½ inches) collected by Jack Shumard on 15 June 1987. The maximum length throughout the range is 57.2 cm (22½ inches) (Boundy, 1995).
Distribution:
 This small snake has a peculiar distribution in the state, in that it is found in all but the four corners. Although records from Oklahoma (Sievert and Sievert, 2005) indicate that it may occur in the southwest corner of the state along the Cimarron River.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
Open icons are questionable records; Click on a marker to view details.
Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 1,365
    Records 
  • 1,299
    Museum Vouchers 
  • 66
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
 County Breakdown: County Name (# occurrences):
 Allen (15); Anderson (7); Atchison (6); Barber (44); Barton (12); Bourbon (2); Brown (6); Butler (15); Chase (57); Chautauqua (8); Clark (14); Clay (6); Cloud (1); Coffey (4); Comanche (2); Cowley (108); Crawford (19); Dickinson (24); Douglas (43); Elk (33); Ellis (31); Ellsworth (18); Finney (24); Ford (4); Franklin (12); Geary (18); Gove (1); Graham (1); Gray (1); Greenwood (8); Hamilton (2); Harper (4); Harvey (7); Hodgeman (5); Jackson (1); Jefferson (4); Jewell (31); Johnson (40); Kearney (1); Kingman (4); Kiowa (7); Labette (1); Lane (3); Leavenworth (25); Lincoln (12); Linn (3); Logan (8); Lyon (13); Marion (2); Marshall (5); McPherson (8); Miami (5); Mitchell (4); Montgomery (3); Morris (7); Nemaha (22); Neosho (7); Ness (3); Norton (1); Osage (9); Osborne (3); Ottawa (18); Phillips (3); Pottawatomie (53); Pratt (9); Reno (12); Republic (3); Rice (4); Riley (26); Rooks (3); Rush (12); Russell (143); Saline (17); Scott (14); Sedgwick (140); Shawnee (36); Smith (1); Stafford (5); Sumner (15); Trego (5); Unknown (10); Wabaunsee (22); Washington (5); Wichita (1); Wilson (8); Woodson (4); Wyandotte (2);

Fossil History:
 Pleistocene fossil specimens are known from McPherson and Rice counties.
Fossils from the Williams Local Fauna of Rice County (Pleistocene: Illinoian) (Preston 1979, Holman 1984; Holman 1995) are assignable to this taxon.
Fossils from the Sandahl Local Fauna of McPherson County (Pleistocene: Illinoian) (Holman 1971; Preston 1979; Holman 1995) are assignable to this taxon.
Natural History:
 This secretive snake inhabits the hillsides of open prairies and woodland edge. In addition, it often is found inside towns and cities beneath debris in vacant lots; habitat disturbance by human activity seems to have little effect on its populations. Grant (1937) observed nine specimens struck by automobiles on the streets of Wichita in October and November. This reptile normally is active from April to October. It is nocturnal and spends the day hiding beneath rocks and debris, including dried cow dung.
During winter, it retires beneath the ground to avoid cold temperatures.
Brumwell (1951) found seven examples of the Lined Snake in a retreat located 45.7 cm (18 inches) beneath the ground in a rocky meadow during May in Leavenworth County. Taggart (1992) collected 47 Lined Snakes under rocks on a hillside in Hodgeman County on 5 May and 72 of these reptiles under flat rocks in three hours of collecting in southern Ellis County on 17 June.
Lined Snakes mate in both spring and fall. Courtship has not been recorded in Kansas. Females give birth to young during August, and the number of young per litter ranges from two to twelve (Fitch, 1985), with an average of six or seven. The young attain sexual maturity during their second season.
The Lined Snake feeds exclusively on earthworms. Remarks: Predators of this small snake include larger snakes, birds, and mammals.
Occurrence Activity:
 Number of Unique Obervations (=days): 206; Range: 05 Feb to 07 Dec
Remarks:
 First reported in 'Kansas' (and described to science) by Hallowell (1857) based on a specimen (ANSP 1857: Holotype) presented by Fort Riley post surgeon William A. Hammond. The earliest existing specimen (KU 1752) was collected in Douglas County during July of 1904. The National Museum of Natural History (Paris) has a specimen collected in "Sud-West Kansas" on 27 November 1894.
This snake can be found in a variety of habitats including thin woods, prairies, dumps, and even urban areas. It is usually encountered by turning cover but is also commonly encountered crossing roads at night.
Burt (1935) reported specimens of this snake taken under flat rocks on prairie ledges two miles southwest of Matfield Green, Chase County, on 9 April 1934; and 1 mile northeast of Otto, Cowley County, on 6 May 1934.
Bibliography:
1856 Hallowell, Edward. Notice of a collection of reptiles from Kansas and Nebraska presented to the Academy of Natural Sciences, by Doctor Hammond, U. S. A. Proceedings of the Academy of Natural Sciences, Philadelphia 8():238-253
Contains reference to twenty-four species collected from 'Kansas', and includes the original description of Microps lineatus (=Tropidoclonion lineatum) from Kansas on page 241.
1860 Cope, Edward D. Notes and descriptions of new and little known species of American reptiles. Proceedings of the Academy of Natural Sciences, Philadelphia 12():339-345
1860 Cope, Edward D. Catalogue of Colubridae in the Museum of the Academy of Natural Sciences of Pailadelphia [sic]. I. Calamarinae. Proceedings of the Academy of Natural Sciences, Philadelphia 12():74-79
1875 Cope, Edward D. Check-list of North American Batrachia and Reptilia; with a systematic list of the higher groups, and an essay on geographical distribution. Based on the specimens contained in the U.S. National Museum. Bulletin of the United States National Museum 1():1-104
The first reference to Anaxyrus fowleri (nomen nudum) (not in Kansas). List the following species from Kansas: Carphophiops vermis (= Carphophis vermis); Ophibolus calligaster (= Lampropeltis calligaster); Ophibolus doliatus and Ophibolus doliatus annulatus (= Lampropeltis gentilis/triangulum); Tropidonotus sipedon erythrogaster (= Nerodia erythrogaster); Opheosaurus ventralis (= Ophisaurus attenuatus); Coluber emoryi (= Pantherophis emoryi); Coluber vulpinus (= Pantherophis ramspotti); Pituophis sayi (= Pituophis catenifer); and Tropidoclonion lineatum
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():112-123
Also listed the Scarlet Snake (Cemophora coccinea) [=Ophibolus doliatus var. coccineus] and Red Salamander (Pseudotriton ruber) [=Spelerpes ruber] from Kansas.
1884 Garman, Samuel. The North American reptiles and batrachians. Bulletin of the Essex Institute 16():1-46
1885 Cragin, Francis W. Recent additions to the list of Kansas reptiles and batrachians, with further notes on species previously reported. Bulletin of the Washburn College Laboratory of Natural History 1(3):100-103
1885 Cragin, Francis W. Second contribution to the herpetology of Kansas, with observations on the Kansas fauna. Transactions of the Kansas Academy of Science 9():136-140
1891 Stejneger, Leonhard. Notes on some North American snakes. Proceedings of the United States National Museum 14(876):501-505
Comments on discrepancies in Cope (1975) regarding the validity of  Lampropeltis annulata in Kansas. States that recently described Tropidoclonion lineatum iowa does not deviate from specimens from Kansas including the type specimen.
1892 Garman, H. A synopsis of the reptiles and amphibians of Illinois. Illinois Laboratory of Natural History Bulletin 3():215-403
Discusses the distribution of Tropidoclonion lineatum (page 277) with respect to Kansas.
1900 Cope, Edward D. The crocodilians, lizards and snakes of North America. Pages 153-1270 in Report of the U. S. National Museum for the Year Ending June 30, 1898 , Washington, D. C. pp.
1903 Branson, Edwin B. Snakes of Kansas Thesis. University of Kansas, Lawrence. 41pp.
Describes all snake species reported from Kansas, considering doubtful those species that the author has not encountered himself. This list contains twenty-nine valid species. Collections examined included State University (KU), State House, Washburn College (WU), Ottawa University, State Normal School (ESU), State Agricultural College (KSU), and several high schools in Kansas. Examined the material available to Mozley (1878) and determined that of the thirty-three species listed, only twenty-three species were valid currently. And that Cragin's (1880) list of thirty-two species included eight specimens on the authority of Mozley that this author could not verify in any collection.
1904 Branson, Edwin B. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430
1909 Hurter, Julius and John K. Strecker, Jr. Amphibians and reptiles of Arkansas. Transactions of the Academy of Science St. Louis 18(2):11-27
1911 Hurter, Julius. Herpetology of Missouri. Transactions of the Academy of Science St. Louis 20(5):59-274
1927 Burt, Charles E. An annotated list of the amphibians and reptiles of Riley County, Kansas. Occasional Papers of the Museum of Zoology, University of Michigan (189):12
Accounts on forty species known from the vicinity of Manhattan, Riley County, Kansas. There have been several scientific names changes since publication, which is understandable... however, some usages cannot be explained by subsequent taxonomic reappraisals(e.g.  Gastrophryne carolinensis for Gastrophryne olivacea). The writer lists Eumeces (=Plestiodon) obsoletus and E. guttulatus yet correctly reasoning that the latter is the young of the former. Within the account of Tantilla gracilis (a common form) the writer mentions that T. nigriceps was reported from Riley County by Branson (1904) but that the specimens at Kansas State were absent at the time of publication. The Prairie Rattlesnake (Crotalus viridis) was first reported from Riley County by Branson (1904) and is reported by the writer based on a specimen in the museum at Kansas State University by Professor E. A. Popenoe. The Prairie Rattlesnake is not currently native to Riley County, and closest reliable localities are 150 to the west.
1929 Taylor, Edward H. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62
1932 Gloyd, Howard K. The herpetological fauna of the Pigeon Lake Region, Miami County, Kansas. Papers of the Museum of Zoology, University of Michigan 15():389-408
First record of Notophthalmus viridescens from Kansas. Second record (after the type locality) of Pseudacris crucifer from Kansas.
1932 Dunn, Emmett R. The status of Tropidoclonion lineatum. Proceedings of the Biological Society of Washington 45():195-198
1933 Stejneger, Leonhard and Thomas Barbour. A Checklist of North American Amphibians and Reptiles. 3rd Edition. Harvard University Press, Cambridge, Massachusetts. pp.
Reference to Kansas is the listed range of several species.
1933 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1935 Burt, Charles E. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16(3):311-366
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1937 Grant, Chapman. Herpetological notes from Central Kansas. American Midland Naturalist 18(3):370-372
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1941 Schmidt, Karl Peterson and D. D. Davis. Field Book of Snakes of the United States and Canada. C.P. Putnam and Sons, New York. 365pp.
1942 Hudson, G. E. The amphibians and reptiles of Nebraska. Nebraska Conservation Bulletin 24():1-146
1945 Bugbee, R. E. A note on the mortality of snakes on highways in western Kansas. Transactions of the Kansas Academy of Science 47():373-374
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1951 Brumwell, Malcolm J. An ecological survey of the Fort Leavenworth Military Reservation American Midland Naturalist 45(1):187-231
Published posthumously. Lieutenant Brumwell died December 14, 1941, as a result of injuries incurred during the Japanese attack on Pearl Harbor. This paper is a condensed version of his thesis for the Master's degree.
1953 Schmidt, Karl P. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280pp.
Schmidt's first edition of his standardized checklist to North American amphibians and reptiles. Includes several specific references to Kansas in the range descriptions.
1953 Ramsey, L. W. The lined snake, Tropidoclonion lineatum (Hallowell). Herpetologica 9(1):7-24
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1958 Smith, Ronald E. Natural history of the Prairie Dog in Kansas. University of Kansas Museum of Natural History, Miscellaneous Publications (16):1-36
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1967 Choate, Jerry R. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46pp.
1971 Holman, J. Alan. Herpetofauna of the sandahl local fauna (Pleistocene: Illinoian) of Kansas. Contributions from the Museum of Paleontology, University of Michigan 23(22):349-355
1973 Knight, James L., Eugene D. Fleharty, and Jerry D. Johnson. Noteworthy records of distribution and habits of some Kansas herptiles. Transactions of the Kansas Academy of Science 75(3):273-275
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1977 Knight, James L. and Joseph T. Collins. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19pp.
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
Notable mentions: Spotted Salamander, Smooth Green Snake
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1979 Martin, Larry D. Survey of fossil vertebrates from east-central Kansas: Kansas River bank stabilization study. U.S. Army Corps of Engineers, Kansas City District. 55pp.
1979 Gray, Peter and Eddie Stegall. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8
1979 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1978. Technical Publication of the State Biological Survery of Kansas 8():56-66
1980 Smith, Keith I. Observations of a Captive Kansas Least Weasel (Mustela nivalis). Thesis. Emporia State University, Emporia, Kansas. 55pp.
1980 Spencer, Dwight. Spencer, D. 1980. Ross Natural History Reservation: the first twenty years, 1959 to 1979. Emporia State University, Emporia, Kansas.. 64pp.
1980 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1979. Technical Publication of the State Biological Survery of Kansas 9():1-11
1981 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1980. Technical Publication of the State Biological Survery of Kansas 10():7-19
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1984 Holman, J. Alan. Herpetofaunas of the Duck Creek and Williams Local Faunas (Pleistocene: Illinoian) of Kansas. Pages 20-38 in Contributions in Quaternary Vertebrate Paleontology: A Volume in Memorial to John E. Guilday. Special Publication Number 8. Carnegie Museum of Natural History, Pittsburgh, Pennsylvania. pp.
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Heinrich, Mark L. Herpetofauna of the Konza Prairie Research Natural Area in the Flint Hills region of Kansas with respect to habitat selection. Thesis. Kansas State University, Manhattan, Kansas. 57pp.
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1984 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26
Invalidated the specimens of Thamnophis sirtalis from Hamilton County (reidentified as T. cyrtopsis; KU 2088) and Wallace County mapped in Collins, 1982.
1984 Collins, Joseph T. New records of fishes, amphibians and reptiles in Kansas for 1984. Kansas Herpetological Society Newsletter (58):14-20
1985 Karch, Olin. Summertime in Emporia --  A short Melodrama in 10 acts. Kansas Herpetological Society Newsletter (61):18-24
1985 Fitch, Henry S. Variation in clutch and litter size in New World reptiles. University of Kansas Museum of Natural History Miscellaneous Publication (76):1-76
1985 Lynch, John D. Annotated checklist of the amphibians and reptiles of Nebraska. Transactions of the Nebraska Academy of Science 13():33-57
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1987 Simmons, John E. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894
1988 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19
1988 Nulton, Michael T. and Michael S. Rush. New county records of amphibians and reptiles in Gray County, Kansas. Kansas Herpetological Society Newsletter (74):10-12
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1988. Kansas Herpetological Society Newsletter (75):15-18
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Fitch, Henry S. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Ecology and Hydrology of Kansas Ecological Reserves and the Baker Wetlands. Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence, Kansas. pp.
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1992 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17
1992 Taggart, Travis W. Observations on Kansas amphibians and reptiles Kansas Herpetological Society Newsletter (88):13-15
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Taggart, Travis W. KHS field trips. Kansas Herpetological Society Newsletter (91):3
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1993 Viets, Brian E. An annotated list of the herpetofauna of the F. B., and Rena G. Ross Natural History Reservation. Transactions of the Kansas Academy of Science 96(1/2):103-113
1993 Fitch, Henry S. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224
1994 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4
See, Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14.
1994 Dloogatch , Michael A. (Editor) Herpetology 1994 Bulletin of the Chicago Herpetological Society 29(12):282-283
Note on the Fitch (1993) paper in the Transactions of the Kansas Academy of Sciences (96(3-4): 213-224) on the abundance.
1994 Smith, Hobart M. and David Chiszar. Variation in the Lined Snake (Tropidoclonion lineatum) in northern Texas. Bulletin of the Maryland Herpetological Society 30(1):6-14
1995 Holman, J. Alan. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243pp.
1995 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1994. Kansas Herpetological Society Newsletter (100):24-47
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1995 Boundy, Jeff. Maximum lengths of North American snakes. Bulletin of the Chicago Herpetological Society 30(6):109-122
1996 Miller, Larry L. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3
1996 Miller, Larry L. Many amphibian and reptile species identified during KHS 1996 fall field trip to Wabaunsee County. Kansas Herpetological Society Newsletter (106):2-3
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1997 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1996. Kansas Herpetological Society Newsletter (107):14-16
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1998 Gamble, Jerre. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1998 Rundquist, Eric M. KDWP herp sting so far nets nine on Kansas and Federal charges. Kansas Herpetological Society Newsletter (112):5-6
1998 Collins, Joseph T. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
2000 Taggart, Travis W. KHS spring field trip sets record for attendance. Kansas Herpetological Society Newsletter (120):5-5
2000 Taggart, Travis W. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2001 Alfaro, Michael E. and Stevan J. Arnold. Molecular systematics and evolution of Regina and the Thamnophiine snakes. Molecular Phylogenetics and Evolution 21(3):408-423
The T. lineatum used in their analyses was from Russell County, Kansas.
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Fogell, Daniel D. Occurrence and relative abundance of amphibians and reptiles at Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. Interim Report. National Park Service, Washington, D.C.. 6pp.
2002 Ellis, Mark R. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5
2002 Taggart, Travis W. Results of the KHS 2002 fall field Trip. Journal of Kansas Herpetology (4):11-13
To Washington County, Kansas (also includes ancillary counts from Clay and Marshall counties).
2002 Gubanyi, James E. Osage County herp count I. Journal of Kansas Herpetology (4):15
2003 Fogell, Daniel D. A herpetofaunal inventory of Tallgrass Prairie National Preserve, Homestead National Monument of America, and Pipestone National Monument within the Heartland Inventory and Monitoring Network. National Park Service, Washington, D.C.. 59pp.
This is the version the author submitted to the NPS. Their final publication was modified.
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Platt, Dwight R. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (7):10
2003 Volkmann, Al. Cowley County herp count 1. Journal of Kansas Herpetology (7):7
2004 Volkmann, Al. Cowley County herp count. Journal of Kansas Herpetology (11):10
2004 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (11):11-12
2004 Taggart, Travis W. Results of the KHS 2004 fall feld trip . Journal of Kansas Herpetology (12):15-16
2005 Taggart, Travis W., Curtis J. Schmidt, and Richard S. Hayes. Geographic distribution: Tropidoclonion lineatum. Journal of Kansas Herpetology (13):10
2005 Taggart, Travis W. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Wilgers, Dustin J. and Eva A. Horne. Effects of different burn regimes on tallgrass prairie herpetofaunal species diversity and community composition in the Flint Hills, Kansas. Journal of Herpetology 40():73-84
2006 Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62():378-388
2007 Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5pp.
2008 Taggart, Travis W. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3
2009 Murrow, Daniel G. KHS 2009 spring field trip. Journal of Kansas Herpetology (29):42769
2009 Schmidt, Curtis J. Geographic distribution: Tropidoclonion lineatum. Journal of Kansas Herpetology (30):11
County record, Marion County, Kansas.
2010 Miller, Larry L. Investigation of the Checkered Garter Snake in Kansas with notes on other Amphibians, Reptiles, and Turtles encountered. Kansas Department of Wildlife and Parks, Pratt, Kansas. 31pp.
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2010 Collins, Suzanne L. and Joseph T. Collins. Geographic Distribution: Tropidoclonion lineatum (Lined Snake). Journal of Kansas Herpetology (33):6
Second record, Ottawa County, Kansas.
2010 Gray, Brian S. Distribution of native and exotic earthworms in the eastern United States: Implications for the ecology of vermivorous snakes. Bulletin of the Chicago Herpetological Society 45(5):73-86
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2011 Houck, Mike. Fort Riley Herpetofaunal Survey for 2011. Journal of Kansas Herpetology (39):9
2011 Shofner, Ryan and Sara Unruh. Geographic Distribution: Tropidoclonion lineatum. Kansas: Finney County. Journal of Kansas Herpetology (40):8
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2012 Hamilton, Bryan Tyler, Rachel Hart , and Jack W. Sites Jr. Feeding ecology of the Milksnake (Lampropeltis triangulum, Colubridae) in the western United States. Journal of Herpetology 46(4):515-522
2013 Sinclair, Tom. A four-day spring snake count across northern Kansas. Collinsorum 2(1/2):9
2013 Baldwin, Mary Kate. “Herps in Havensville” Herp Surveys. Collinsorum 2(3/4):10
2013 Taggart, Travis W. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2015 Brown, Kasandra A. Occupancy Modeling Of Herpetofauna And Grassland Nesting Birds At Quivira National Wildlife Refuge. Thesis. Fort Hays State University, Hays, Kansas. 72pp.
2015 Mike Houck. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102
2017 Mardis, Dexter R. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10
2017 Taggart, Travis W. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8
2017 Schmidt, Curtis J. Herp Count: Southeast Ellis County. Collinsorum 6(2-3):9
2017 Taggart, Travis W. Herp Count: Clark County State Lake. Collinsorum 6(2-3):9
2017 Taggart, Travis W. Herp Count: Northeast Barton County. Collinsorum 6(2-3):9
2018 Houck, Mike. Herp Count: Fort Riley Military Installation Collinsorum 7(1):17
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2021 Schmidt, Curtis J. Herp Count: Ellis County: KHS-2020-32. Collinsorum 9(3):16
2023 Russell, Elisabeth Habitat associations and fine-scale movements of the Red-spotted Toad (Anaxyrus punctatus) in Kansas and the efficacy of remote telemetry for monitoring small-scale movements. Thesis. Fort Hays State University, Hays, Kansas. 81pp.
2023 Dunbar, Oliver L., Keith Geluso, and Daniel D. Fogell. Geographic distribution: Tropidoclonion lineatum. USA, Nebraska. Herpetological Review 54(4):603
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Travis W. Taggart © 1999-2024 — w/ Sternberg Museum of Natural History, Fort Hays State University