An adult Rough Greensnake from Allen County. Image by Lisa Wehrly.
An adult Rough Greensnake from Bourbon County, Kansas. Image © Dan Fogell.
An adult Rough Greensnake from Allen County. Image by Lisa Wehrly.
REPTILIA (Reptiles) SQUAMATA (PART) (Snakes) COLUBRIDAE (Harmless Egg-laying Snakes)

Rough Greensnake
Opheodrys aestivus (Linnaeus 1766)
ō-fē-ō-drēz — ĕs-tĭ-vŭs


Conservation Status:

State: None

Federal: None
NatureServe State: S4 - Apparently Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
HARMLESS. The Rough Greensnake is characterized by keeled scales, a divided anal scale, and a bright green head, body, and tail. The belly is a uniform white or cream, and the chin may be light yellow. Adult males have a slightly longer tail than females. The tongue is red and slightly darker at the fork tips.
Adults normally grow 56.0-81.0 cm (22-32 inches) in total length. The largest specimen from Kansas is a female (KU 45334) from Elk County with a total length of 87.7 cm (34½ inches) collected by Henry S. Fitch on 8 July 1957. The maximum length throughout the range is 114.9 cm (45¼ inches) (Powell et al., 2016).

Distribution:
Known from moderately to heavily wooded areas in southeast Kansas from the Arkansas River drainage north to the Kansas River basin.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
Open icons are questionable records; Click on a marker to view details.
Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 591
    Records 
  • 565
    Museum Vouchers 
  • 26
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (88); Anderson (16); Bourbon (30); Butler (7); Chautauqua (36); Cherokee (45); Cowley (19); Crawford (40); Douglas (2); Elk (22); Franklin (30); Greenwood (9); Johnson (93); Labette (56); Linn (11); Miami (16); Montgomery (21); Neosho (18); Sumner (2); Wilson (15); Woodson (9); Wyandotte (6);

Fossil History:
Not known from Kansas.

Natural History:
This slender, graceful snake inhabits trees, brush, shrubbery, and bushes near aquatic situations, particularly small lakes. Jeffrey Burkhart (pers. comm., 1979; Collins, 1993) found two Rough Greensnakes along a stream in Cherokee County on 21 July. Collins (1982) reported a single example of this snake on a branch overhanging a stream in the same county on 9 April. When not gliding through vegetation in search of food, this snake retires beneath rocks or leaves, or in hollow logs. The Rough Greensnake blends perfectly with green plants and is extremely difficult to observe. Burt and Hoyle (1935) stated that they may be found under rocks in prairie ledges in the early spring, but later they are most likely to be located in vegetation, especially in shrubs and trees.
This snake exhibits another interesting adaptation for concealment in thick vegetation. When a breeze causes the stems of plants to bend and wave, a Rough Greensnake sitting in their midst will bend and wave in unison with the vegetation.
This snake is active from April to September, is strictly diurnal, and has a small home range. At night it sleeps in shrubs and bushes. During winter, the Rough Greensnake retreats deep into leaf piles, in burrows beneath the ground, or in hollow logs for warmth.
The Rough Greensnake mates during the spring, but courtship is not known in Kansas. During June or July, a female lays a clutch of eggs, which range in number from one to ten (Fitch, 1985), with an average of five or six. Robert F. Clarke (pers. comm., 1980; Collins, 1993) obtained an adult female Rough Greensnake from Woodson County on 3 July. The snake laid ten eggs on 12-13 July. The eggs of this reptile may hatch in two months.
Rough Greensnakes are primarily insect-eaters, readily consuming grasshoppers, caterpillars, and crickets, but will also eat spiders. These reptiles are active daytime predators that locate prey by sight, slowly approach it, and grab it with a short rapid strike.
Taggart (1992) observed two Rough Greensnakes being consumed by a Racer on 8 September in Woodson County.

Occurrence Activity:
Number of Unique Obervations (=days): 143; Range: 05 Feb to 25 Nov
Remarks:
First reported from Kansas by Cragin (1885) based on a specimen collected by Colonel Nathaniel S. Goss at Neosho Falls (Woodson County). The earliest specimens (KU 1578-83: Bourbon County; KU 2701: Franklin County; KU 2702-3; Labette County) from Kansas were all collected during June of 1911.
This small, slender snake blends in well with shrubs, trees, and grasses it spends most of its time in. Most encounters with this species occur as it attempts to cross roadways In the spring, they are often found under rocks or another surface cover.
Based on a captive specimen, Snider and Bowler (1992) reported a maximum longevity for this snake of six years, eight months, and 21 days.

Bibliography:
1766 Linné, Carl von (=Linneaus). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tomus I. [The system of nature through the three kingdoms of nature, according to classes, orders, genera, species with characters, differences, synonyms, places. Volume I.] Laurentii Salvii, Stockholm, Holmiae, Editio duodecima, reformata (12th Edition). 1-532pp.
Contains the original descriptions of Lacerta 6-lineata (=Aspidoscelis sexlineata) page 364, Sistrurus miliarius page 372, Boa contortrix (=Agkistrodon contortrix): page 373, Coluber striatulus (=Haldea striatula) page 275, Coluber punctatus (=Diadophis punctatus) page 376, and Coluber aestivus (=Opheodrys aestivus) page 387.
1885 Cragin, Francis W. Recent additions to the list of Kansas reptiles and batrachians, with further notes on species previously reported. Bulletin of the Washburn College Laboratory of Natural History 1(3):100-103
1885 Cragin, Francis W. On the Washburn Biological Survey of Kansas. Kansas City Review of Science 8(10):576-580
First report of the Rough Green Snake (Opheodrys aestivus) in Kansas, collected by Col. N. S. Goss from Neosho Falls, Woodson, County.
1885 Cragin, Francis W. Second contribution to the herpetology of Kansas, with observations on the Kansas fauna. Transactions of the Kansas Academy of Science 9():136-140
1904 Branson, Edwin B. Snakes of Kansas. University of Kansas Science Bulletin 2(13):353-430
1911 Hurter, Julius. Herpetology of Missouri. Transactions of the Academy of Science St. Louis 20(5):59-274
1922 Loding, H. P. A preliminary catalogue of Alabama amphibians and reptiles. Geological Survey of Alabama, Museum Paper (5):59
1929 Taylor, Edward H. A revised checklist of the snakes of Kansas. University of Kansas Science Bulletin 19(5):53-62
1931 Haltom, William L. Alabama reptiles. Alabama Museum of Natural History Museum Paper (11):1-145
1933 Stejneger, Leonhard and Thomas Barbour. A Checklist of North American Amphibians and Reptiles. 3rd Edition. Harvard University Press, Cambridge, Massachusetts. pp.
Reference to Kansas is the listed range of several species.
1933 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1936 Hurd, Myron Alec. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103pp.
Under the supervision of thesis adviser Harry H. Hall. Report on 38 species (8 turtles, 7 lizards, and 23 snakes)... most unsubstantiated. Interesting inclusion are Crotalus horridus, Crotalus viridis, Kinosternon subrubrum, Opheodrys vernalis, and Phrynosoma cornutum.
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1941 Schmidt, Karl Peterson and D. D. Davis. Field Book of Snakes of the United States and Canada. C.P. Putnam and Sons, New York. 365pp.
1941 Mansueti, Romeo. A descriptive catalogue of the amphibians and reptiles found in and around Baltimore City, Maryland, within a radius of twenty miles. Proceedings of the Natural History Society of Maryland 7():1-53
1947 Hall, Henry H. and Hobart M. Smith. Selected records of reptiles and amphibians from southeastern Kansas Transactions of the Kansas Academy of Science 49(4):447-454
Report on certain Kansas specimens housed in the collection at what is now Pittsburg State University. Included are several species of dubious status today, including Cryptobranchus alleganiensis from the Neosho and Spring rivers (the only specimens from those significant drainages ever documented), Ambystoma maculatum from just north of Pittsburg, Crawford County, Heterodon nasicus from Crawford County, Opheodrys vernalis from Crawford County, Sonora episcopa from Crawford County, Agkistrodon piscivorus from Cherokee County, Crotalus atrox from Crawford County, and Crotalus viridis from Crawford County. They report several significant range extensions including Kinosternon flavescens from Turkey Creek in southeast Cherokee County, Graptemys geographica from just north of Pittsburg, Crotaphytus collaris from near Columbus, Cherokee County, Sceloporus consobrinus from just north of Pittsburg, Phrynosoma cornutum from Cherokee and Crawford counties, Heterodon platirhinos from Cherokee and Crawford counties, Haldea striatula from Crawford County, Sistrurus tergeminus from Crawford County, and a 402 lb Macrochelys temminckii in Cherokee County from just east of Chetopa (Labette County). They allude to the potential for Anaxyrus fowleri to occur in southeast Kansas and for native populations of Crotalus atrox in south central Kansas (in part from the disclosure that John R. Breukelman [then of ESU] had obtained three specimens in Woods County Oklahoma, 3/4 of a mile south of the Kansas line). None of the specimens the paper was based on, exist today.
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1953 Schmidt, Karl P. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280pp.
Schmidt's first edition of his standardized checklist to North American amphibians and reptiles. Includes several specific references to Kansas in the range descriptions.
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1967 Choate, Jerry R. Wildlife in the Wakarusa Watershed of Northeastern Kansas. Kansas Biological Survey, Lawrence. 46pp.
1967 Gier, Herschel T. Vertebrates of the Flint Hills. Transactions of the Kansas Academy of Science 70(1):51-59
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1977 Perry, Janice. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
Notable mentions: Spotted Salamander, Smooth Green Snake
1979 Gray, Peter. Low attendance slows KHS. Kansas Herpetological Society Newsletter (32):1
1982 Collins, Joseph T. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57pp.
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1984 Grobman, Arnold B. Scutellation variation in Opheodrys aestivus. Bulletin of the Florida State Museum, Biological Sciences 29(3):153-170
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1992 Taggart, Travis W. Observations on Kansas amphibians and reptiles Kansas Herpetological Society Newsletter (88):13-15
1992 Grobman, Arnold B. On races, clines, and common names in Opheodrys. Herpetological Review 23(1):14-15
1992 Grobman, Arnold B. Metamerism in the snake Opheodrys vernalis, with a description of a new subspecies. Journal of Herpetology 26():175-186
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Fitch, Henry S. Relative abundance of snakes in Kansas. Transactions of the Kansas Academy of Science 96(3/4):213-224
1994 Fitzgerald, Eve C. and Charles Nilon. Classification of habitats for endangered and threatened species in Wyandotte County, Kansas Kansas Department of Wildlife and Parks, Pratt, Kansas. 98pp.
1994 Riedle, J. Daren. Distribution of the Timber Rattlesnake (Crotalus horridus) in Chautauqua, Elk, and Montgomery counties, Kansas. Kansas Herpetological Society Newsletter (95):43051
1994 Riedle, J. Daren. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Boundy, Jeff. Maximum lengths of North American snakes. Bulletin of the Chicago Herpetological Society 30(6):109-122
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1997 Collins, Joseph T. A report on the KHS fall field trip to the Marais des Cygnes wildlife refuges. Kansas Herpetological Society Newsletter (110):2-3
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1998 Gamble, Jerre. Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. KDWP herp sting so far nets nine on Kansas and Federal charges. Kansas Herpetological Society Newsletter (112):5-6
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
1999 Taggart, Travis W. Cherokee County fall 1999 herp count. Kansas Herpetological Society Newsletter (117):6
Reported Anaxyrus woodhousii was likely A. fowleri.
2000 Taggart, Travis W. Results of the KHS 2000 fall field trip. Kansas Herpetological Society Newsletter (122):6-8
2000 Walley, Harlan D. and Michael V. Plummer. Opheodrys aestivus. Catalogue of American Amphibians and Reptiles (718):1-14
2001 Taggart, Travis W. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14
2001 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 2000. Kansas Herpetological Society Newsletter (124):6-8
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Riedle, J. Daren and A. Hynek. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20
2003 Taggart, Travis W. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12
2003 Platt, Dwight R. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2005 Taggart, Travis W. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the KHS Spring Field Trip to Chautauqua County. Journal of Kansas Herpetology (38):2-4
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2013 Miller, Larry L. and Suzanne L. . Range extension of the Rough Green Snake (Opheodrys aestivus) in Kansas. Collinsorum 2(1/2):12
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4
2013 Taggart, Travis W. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2014 Taggart, Travis W. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12
2014 Taggart, Travis W. Recent scientific and standard English name changes effecting the Kansas herpetofauna. Collinsorum 3(2-4):9-10
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Mardis, Dexter R. Geographic distribution: Opheodrys aestivus: Elk County, Kansas. Collinsorum 5(4):8-17
2016 Taggart, Travis W. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102
2017 Mardis, Dexter R. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10
2017 Taggart, Travis W. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8
2018 Konvalina, John D. and Stanley E. Trauth. Seasonal variation of testicular tissue in Northern Rough Greensnakes, Opheodrys a. aestivus, from Alabama. Southeastern Naturalist 17(3):521-530
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2020 Hullinger, Allison, Zackary Cordes, Daren Riedle, and William Stark. Habitat assessment of the Broad-headed Skink (Plestiodon laticeps) and the associated squamate community in eastern Kansas. Transactions of the Kansas Academy of Science 123(1-2):137-150
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Neosho County: KHS-2020-01 Collinsorum 9(3):11
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Cherokee County: KHS-2020-02 Collinsorum 9(3):11-12
2021 Rittmeyer, Eric N., Ahmed Elmonier, and Sara Ruane. Phylogeography of the Rough Greensnake, Opheodrys aestivus (Squamata: Colubridae), using multilocus Sanger sequence and genomic ddRADseq data. Journal of Herpetology 55(4):342-354
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Travis W. Taggart © 1999-2024 — w/ Sternberg Museum of Natural History, Fort Hays State University