REPTILIA (Reptiles) LACERTILIA (Lizards) TEIIDAE (Racerunners and Whiptails)

SIX-LINED RACERUNNER
Aspidoscelis sexlineatus (Linnaeus 1766)
ăs-spĭd-ō-skĕl-ĭs — sĕx-lĭn-ē-ā-tŭs


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None

An adult Six-lined Racerunner from Cherokee County, Kansas. © Travis W. Taggart.
An adult Six-lined Racerunner from Russell County, Kansas. © Travis W. Taggart.
Image © Suzanne L. Collins, CNAH.

Diagnosis:
This colorful lizard has six light cream, yellow, or green stripes running from the neck onto the tail (three on each side). Many specimens will have a seventh stripe running down the middle of the back, although when this stripe is present, it is seldom as distinct as the more lateral stripes. The background color is tan to dark brown on the back and becomes bright green along the sides, especially in warm adult males, which often also have bright, light blue jaws that may extend onto the throat and belly. The scales on the back are small and granular. The belly scales are much larger and rectangular in shape.
Adults normally 152-203 mm (6-8 inches) in total length. The largest specimen from Kansas is a male (KU 220236) from Cherokee County with a snout-vent length of 82 mm and a total length of 233 mm (9 1/8 inches) collected by Randall Reiserer on 18 May 1992. The maximum length throughout the range is 10½ inches (Conant and Collins, 1998).

Distribution:
This lizard occurs throughout Kansas, and may particularly abundant in areas characterized by loose soils and patchy open (or rocky) vegetation. A quick and alert species, they are most easily taken under cover early in the morning.
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  • Occurrence Summary:  
  • 1,774
    Records 
  • 1,556
    Museum Vouchers 
  • 218
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (25); Anderson (19); Barber (32); Barton (8); Bourbon (25); Butler (1); Chase (1); Chautauqua (24); Cherokee (23); Cheyenne (26); Clark (9); Clay (2); Cloud (1); Coffey (3); Comanche (22); Cowley (89); Crawford (47); Decatur (8); Dickinson (4); Doniphan (1); Douglas (31); Edwards (6); Elk (11); Ellis (269); Ellsworth (28); Finney (68); Ford (25); Franklin (4); Geary (8); Gove (28); Graham (3); Grant (5); Gray (18); Greeley (1); Greenwood (30); Hamilton (10); Harper (15); Harvey (8); Haskell (1); Hodgeman (19); Jefferson (2); Jewell (2); Johnson (3); Kearney (2); Kingman (48); Kiowa (17); Labette (11); Lane (2); Leavenworth (3); Lincoln (11); Linn (2); Logan (18); Lyon (5); McPherson (3); Meade (44); Miami (2); Mitchell (6); Montgomery (12); Morton (38); Neosho (16); Ness (2); Norton (7); Osage (2); Osborne (4); Ottawa (7); Pawnee (5); Phillips (9); Pottawatomie (16); Pratt (19); Rawlins (11); Reno (30); Republic (6); Rice (9); Riley (50); Rooks (8); Rush (2); Russell (130); Saline (19); Scott (12); Sedgwick (30); Seward (10); Shawnee (8); Sheridan (2); Sherman (2); Smith (1); Stafford (10); Stanton (5); Stevens (3); Sumner (5); Thomas (1); Trego (54); Unknown (11); Wabaunsee (3); Wallace (4); Washington (34); Wichita (1); Wilson (33); Woodson (2); Wyandotte (2);

Natural History:
As their name implies, they are quick and are usually just a blur when you encounter them. These lizards are easiest to observe by turning cover on cool mornings. They are most common in areas with sandy soils.
By the heat of the day, they are out actively searching for invertebrates to eat. Hartman (1906) summarized that due to their speed, no insect could escape them, except by flight. He reported grasshoppers, cockroaches, tiger-beetles, and other beetles as prey items.

Occurrence Activity:
Remarks:
First reported in Kansas by Hallowell (1857) based on a specimen procured by Fort Riley surgeon William A. Hammond in the vicinity of Fort Riley. Bocourt (1879) gives the following account (page 278; translated from French) "The Museum [National Museum of Natural History (Paris)] has nine copies of this variety, which, according to the method of coloration, show notable differences: the smallest, from Kansas, was donated as Cnemidophorus gularis by the Academy of Philadelphia; his body is crossed by six yellow longitudinal lines; the interval between them is decorated with small black circular lines, the lower regions are of a pale yellow." The earliest existing specimen (MCZ 2458) was collected from "Leavenworth or Topeka, Kan" sometime in 1871.
Cragin (1885) reported a specimen from McPherson County (as Cnemidophornns tessellatus [=Aspidoscelis tessellatus] in error) collected by Dr. John Rundstrom.

Bibliography:
1766 Linné, Carl von (=Linneaus). Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species cum characteribus, differentiis, synonymis, locis. Tomus I Laurentii Salvii, Stockholm, Holmiae, Editio duodecima, reformata (12th Edition). 1-532pp.
Contains the original descriptions of Lacerta 6-lineata (=Aspidoscelis sexlineata) page 364, Sistrurus miliarius page 372, Boa contortrix (=Agkistrodon contortrix): page 373, Coluber striatulus (=Haldea striatula) page 275, Coluber punctatus (=Diadophis punctatus) page 376, and Coluber aestivus (=Opheodrys aestivus) page 387.
1848 Abert, James W. Notes of Lieutenant J. W. Abert. Appendix Number 6. Pages 386-414 in Notes of a Military Reconnaissance from Fort Leavenworth, in Missouri to San Diego, in California including Parts of Arkansas, Del Norte and Gila Rivers Wendell and Benthuysen, Printers, Senate Executive Document 41, 30th Congress First Session, Washington, D. C. pp.
Lieutenant Abert left Fort Leavenworth on 27 June 1846 en route to San Diego, California.
By 8 July they had reached Turkey Creek in Marion County, where he remarked "The men killed several rattlesnakes near our camp, and one a grey snake, marked with a row of blackish spots along the back; it is said never to exceed two feet in length, and is called the grey rattlesnake."
On 11 July they had reach Cow Creek in Rice County and Abert noted "As our wagons moved along the road, the lizards (Lacerta lineatus) [Aspidoscelis sexlineata] were darting rapidly along the ruts in front of it, anxious to escape being crushed. The common land turtle (Testudo clausa) [Terrepene ornata] were also very abundant."
On 14 July they had reached Pawnee Rock (Barton County) where he wrote, "This morning Laing brought me a very large toad, (Rana musica) [currently a synonym for Anaxyrus terrestris but likely referring to A. woodhousii far exceeding any I ever before have seen."
On 15 July, and still at Pawnee Rock he wrote "To-day we saw several large white cranes with black-tipped wings; (Grus americanus) and Laing killed me some rattlesnakes, (Crotalus horridus) [not local] and several prairie snakes."
On 18 July they were moving along the Arkansas River west of Pawnee Rock and noted "The ruts of the road were full of little lizards, sunning themselves; as we approached they would dart briskly away, manifestly disinclined to play the part of devotees to Juggernaut."
1857 Hallowell, Edward. Notice of a collection of reptiles from Kansas and Nebraska presented to the Academy of Natural Sciences, by Doctor Hammond, U. S. A. Proceedings of the Academy of Natural Sciences, Philadelphia 8():238-253
Contains reference to twenty-four species collected from 'Kansas', and includes the original description of Microps lineatus (=Tropidoclonion lineatum) from Kansas on page 241.
1857 Hallowell, Edward. Note on the collection of reptiles from the neighborhood of San Antonio, Texas, recently presented to the Academy of Natural Sciences by Dr. A. Heerman. Proceedings of the Academy of Natural Sciences, Philadelphia 8():306-310
Report on a lot of specimens received at the Museum of the Academy of Natural Sciences from Dr. Hammond of Ft. Riley Kansas. Of special note are three specimens of Phyrnosoma douglassii (=Phyrnosoma hernandesi) from Ft Riley (it does not naturally occur there and likely never did). Several of the specimens are marked Bridger's Pass (in Wyoming) (Western Milksnake, Plains Hog-nosed Snake, Slender Glass Lizard, and Western Tiger Salamander; all but the Slender Glass Lizard occur near there). A specimen of the Red-spotted Toad is reported as well, however, it does not occur near Ft. Riley, and as given "One Bufo punctatus, (young of Americanus)." there is some question as to its actual identity.
1879 Bocourt, M. F. Etudes sur les reptiles, p. i-xiv, 1-1012. In Recherches Zoologiques pour servir a l'Histoire de Ia Faune de l'Amérique Centrale et du Mexique. Mission Scientifique au Mexique et dans l'Amérique Centrale, Recherches zoologiques. Part 2, sect. 1; In A Imprimerie Impériale, Paris [3, Pt. 6]. A Imprimerie Impériale, Paris, France.. 360–440pp.
In French. The original description of Plestiodon septentrionalis obtusirostris, (based on a specimen from Texas); illustrations of a Plestiodon obsoletus and a Coluber constrictor from Kansas. In text comments on Phrynosoma hernandesi (as Tapaya brevirostris), Aspidoscelis sexlineata, Plestiodon obsoletus, Coluber constrictor, and Heterodon nasicus from Kansas.
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():112-123
Also listed the Scarlet Snake (Cemophora coccinea) [=Ophibolus doliatus var. coccineus] and Red Salamander (Pseudotriton ruber) [=Spelerpes ruber] from Kansas.
1885 Cragin, Francis W. Recent additions to the list of Kansas reptiles and batrachians, with further notes on species previously reported. Bulletin of the Washburn College Laboratory of Natural History 1(3):100-103
1885 Cragin, Francis W. Second contribution to the herpetology of Kansas, with observations on the Kansas fauna. Transactions of the Kansas Academy of Science 9():136-140
1900 Cope, Edward D. The crocodilians, lizards and snakes of North America. Pages 153-1270 in Report of the U. S. National Museum for the Year Ending June 30, 1898 , Washington, D. C. pp.
1906 Hartman, Frank A. Food habits of Kansas lizards and batrachians. Transactions of the Kansas Academy of Science 20():225-229
1916 Householder, Victor H. The Lizards and Turtles of Kansas with Notes on Their Distribution and Habitat. Thesis. University of Kansas, Lawrence. 100pp.
1920 Taylor, Edward H. The Lizards of Kansas with Notes on Habits. Thesis. University of Kansas, Lawrence. 117pp.
Though Dr. Taylor's thesis lists 1916 as the publication date (as does version that the KHS published in 1993). His degree was not awarded until 1920, which is the official publication date.
1928 Burt, Charles E. A key to the species of lizards definitely reported from Kansas. Privately printed, Enterprise Press, Bristow, Nebraska. pp.
Essentially a separate from the writer's "Lizards of. Kansas" which was in-press in the Transactions of the Academy of Science of St. Louis (Burt 1928. 26(1):1-81). Includes a glossary and a note on the variation in Sceloporus undulatus thayerii (= Sceloporus consobrinus).
1928 Burt, Charles E. Insect food of Kansas lizards with notes on feeding habits. Journal of the Kansas Entomological Society 1(3):50-68
1929 Taylor, Edward H. List of reptiles and batrachians of Morton County, Kansas, reporting species new to the state fauna. University of Kansas Science Bulletin 19(6):63-65
Annotated listing of 25 species discovered in southwest Morton County just prior to the Dust Bowl that wreaked havoc on the region and the subsequent creation of the Cimarron National Grassland. Of special note are Heterodon platirhinos, Thamnophis marcianus, and Anaxyrus debilis (all of which have not been found in the area since).
1933 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error.
1933 Burt, Charles E. Some distributional and ecological records of Kansas reptiles. Transactions of the Academy of Science St. Louis 26():186-208
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1934 Burt, Charles E. and W. L. Hoyle. Additional records of the reptiles of the central prairie region of the United States. Transactions of the Kansas Academy of Science 37():193-216
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1935 Burt, Charles E. Further records of the ecology and distribution of amphibians and reptiles in the middle west. American Midland Naturalist 16(3):311-366
1935 Burt, Charles E. and William L. Hoyle. Additional records of the reptiles of the central prairie region of the United States Transactions of the Kansas Academy of Science 37():193-216
1936 Hurd, Myron Alec. The reptiles of Cherokee County, Kansas. Thesis. Pittsburg State University, Pittsburg, Kansas. 103pp.
Under the supervision of thesis adviser Harry H. Hall. Report on 38 species (8 turtles, 7 lizards, and 23 snakes)... most unsubstantiated. Interesting inclusion are Crotalus horridus, Crotalus viridis, Kinosternon subrubrum, Opheodrys vernalis, and Phrynosoma cornutum.
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1939 Tihen, Joseph A. and James M. Sprague Amphibians, reptiles, and mammals of the Meade County State Park Transactions of the Kansas Academy of Science 42():499-512
1940 Taylor, Edward H. Palatal sesamoid bones and palatal teeth in Cnemidophorus, with notes on these teeth in other saurian genera. Proceedings of the Biological Society of Washington 53():119-124
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1947 Hall, Henry H. and Hobart M. Smith Selected records of reptiles and amphibians from southeastern Kansas Transactions of the Kansas Academy of Science 49(4):447-454
Report on certain Kansas specimens housed in the collection at what is now Pittsburg State University. Included are several species of dubious status today, including Cryptobranchus alleganiensis from the Neosho and Spring rivers (the only specimens from those significant drainages ever documented), Ambystoma maculatum from just north of Pittsburg, Crawford County, Heterodon nasicus from Crawford County, Opheodrys vernalis from Crawford County, Sonora episcopa from Crawford County, Agkistrodon piscivorus from Cherokee County, Crotalus atrox from Crawford County, and Crotalus viridis from Crawford County. They report several significant range extensions including Kinosternon flavescens from Turkey Creek in southeast Cherokee County, Graptemys geographica from just north of Pittsburg, Crotaphytus collaris from near Columbus, Cherokee County, Sceloporus consobrinus from just north of Pittsburg, Phrynosoma cornutum from Cherokee and Crawford counties, Heterodon platirhinos from Cherokee and Crawford counties, Haldea striatula from Crawford County, Sistrurus tergeminus from Crawford County, and a 402 lb Macrochelys temminckii in Cherokee County from just east of Chetopa (Labette County). They allude to the potential for Anaxyrus fowleri to occur in southeast Kansas and for native populations of Crotalus atrox in south central Kansas (in part from the disclosure that John R. Breukelman [then of ESU] had obtained three specimens in Woods County Oklahoma, 3/4 of a mile south of the Kansas line). None of the specimens the paper was based on, exist today.
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1951 Wolfenbarger, Keith. A. Systematic and Biological Studies on North American Chiggers of the genus Eutrombicula (Acarina, Trombiculidae). Thesis. University of Kansas, Lawrence. 77pp.
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 species for chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1958 Fitch, Henry S. Natural history of the Six-lined Racerunner (Cnemidophorus sexlineatus). University of Kansas Publications Museum of Natural History 11(2):11-62
1960 Etheridge, Richard E. Additional notes on the lizards of the Cragin Quarry fauna. Papers of the Michigan Academy of Science, Arts and Letters 45():113-117
1960 Etheridge, Richard E. The Pliocene lizard genus Eumecoides Taylor. Bulletin of the Southern California Academy of Sciences 59(2):62-69
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1962 Duellman, William E. and Richard G. Zweifel. A synopsis of the lizards of the sexlineatus group (Genus Cnemidophorus). Bulletin of the American Museum of Natural History 123(3):155-210
1965 Clarke, Robert F. Lizards in Kansas. Kansas School Naturalist 11(4):1-16
1966 Lowe, Charles H. The prairie lined racerunner. Journal of the Arizona Academy of Science 4(1):44-45
Original description of Aspidoscelis sexlineata viridis.
1970 Webb, Robert G. Reptiles of Oklahoma. Stovall Museum, University of Oklahoma Press, Norman. 370pp.
1970 Fitch, Henry S. Reproductive cycles in lizards and snakes. University of Kansas Museum of Natural History Miscellaneous Publication (52):1-247
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Karns, Daryl, Ray E. Ashton, Jr., and Thomas Swearingen. Illustrated Guide to Amphibians and Reptiles in Kansas: An Identification Manual. University of Kansas Publications Museum of Natural History Public Education Series(2):viii + 18
1975 Rundquist, Eric M. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3pp.
Short accounts for twenty-nine recognized amphibians and reptiles from Kingman County, Kansas. With habitat descriptions and for some species, estimates of population density.
1975 Holman, J. Alan. Herpetofauna of the WaKeeney local fauna (Lower Pliocene: Clarendionian) of Trego County, Kansas. Papers on Paleontology, University of Michigan 1974(12):49-66
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1976 Caldwell, Janalee P. and Gregory. Glass. Vertebrates of the Woodson County State Fishing Lake and Game Management Area. Pages 62-76 in Preliminary inventory of the biota of Woodson County State Fishing Lake and Game Management Area. Report No. 5. State Biological Survey of Kansas, Lawrence. pp.
1976 Grow, David. The KHS goes to Chetopa. Kansas Herpetological Society Newsletter (13):2-3
Spring field trip (22 May 1976) along the Neosho River.
1977 Knight, James L. and Joseph T. Collins. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19pp.
1977 Perry, Janice. KHS members achieve goal: Get Cottonmouth. Kansas Herpetological Society Newsletter (21):3-4
1978 Warner, M. and R. Wencel. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16
1978 Schwaner, Terry D. KHS field trip to Grant County, Kansas, 12-14 May 1978. Kansas Herpetological Society Newsletter (25):3-4
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilisThamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1982 Collins, Joseph T. Report to the Kansas Fish and Game Commission on the status of three amphibians in southeastern Kansas. Kansas Fish and Game Commission, Pratt. 57pp.
1982 Fitch, Henry S. Resources of a snake community in prairie-woodland habitat of northeastern Kansas. Pages 83-97 in Herpetological communities: A symposium of the Society for the Study of Amphibians and Reptiles and the Herpetologists League, August 1977.  Wildlife Research Reports 12. 239 pp. U. S. Fish and Wildlife Service, Washington, D. C. pp.
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1983 Trott, Gene. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (52):3-4
1983 Fraser, John C. A trip to the 'TransPecos'. Kansas Herpetological Society Newsletter (54):18-23
1983 Miller, Larry L. Bourbon County field trip well attended and successful. Kansas Herpetological Society Newsletter (54):6-7
1983 Crampton, L. Herpetological collecting in Sumner County, Kansas. Kansas Herpetological Society Newsletter (54):8-9
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1984 Brown, Kenneth L. Pomona: A plains village variant in eastern Kansas and western Missouri. Dissertation. University of Kansas, Lawrence, Kansas. 519pp.
1984 Schwarting, Nancy. KHS field trip, May 1984. Kansas Herpetological Society Newsletter (57):3-4
1985 Miller, Larry L. KHS 1985 field trip to Kirwin Reservoir. Kansas Herpetological Society Newsletter (61):11-12
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1986. Kansas Herpetological Society Newsletter (66):9-16
1987 Simmons, John E. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894
1988 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19
1988 Miller, Larry L. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1988. Kansas Herpetological Society Newsletter (75):15-18
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1989. Kansas Herpetological Society Newsletter (78):16-21
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Collins, Joseph T. and Suzanne L. Collins. Reptiles and Amphibians of the Cimarron National Grasslands, Morton County, Kansas. U. S. Forest Service, Elkhart, Kansas. 60pp.
1991 Fitch, Henry S. Reptiles and amphibians of the Kansas ecological reserves. Pages 71-74 in Multidisciplinary Guidebook 4. Kansas Academy of Science, Lawrence. pp.
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Gubanyi, James E. An observation on the stomach contents of a Texas Longnose Snake (Rhinocheilus lecontei tessellatus). Kansas Herpetological Society Newsletter (89):17
1992 Taggart, Travis W. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4
1992 Rundquist, Eric M. Results of the KHS 1992 fall field trip. Kansas Herpetological Society Newsletter (90):4
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1994 Riedle, J. Daren. A survey of reptiles and amphibians at Montgomery County State Fishing Lake. Kansas Herpetological Society Newsletter (98):11-13
1995 Anderson, Lewis, Mark Shaw, Jeff Blodig, and Tom Walker. Report to the Kansas Department of Wildlife and Parks: Herps encountered during REmap project, summer 1994. Kansas Herpetological Society Newsletter (99):10-17
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1996 Miller, Larry L. Results of the KHS 1995 fall field trip. Kansas Herpetological Society Newsletter (103):3
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Trauth, Stanley E. and Chris T. McAllister. Cnemidophorus sexlineatus. Catalogue of American Amphibians and Reptiles (628):1-12
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1997 Rundquist, Eric M. Addendum to 1997 KHS herp counts. Kansas Herpetological Society Newsletter (109):14-15
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Gamble, Jerre Marais des Cygnes National Wildlife Refuge Comprehensive Conservation Plan U.S. Fish and Wildlife Service, Hartford, Kansas. 91pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1998 Rundquist, Eric M. KDWP herp sting so far nets nine on Kansas and Federal charges. Kansas Herpetological Society Newsletter (112):5-6
1998 Collins, Joseph T. Results of the KHS silver anniversary fall field trip. Kansas Herpetological Society Newsletter (114):6-
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
2000 Van Doren, Mark D. and Curtis J. Schmidt. A herpetological survey of the Fort Larned National Historic Site, Pawnee County, Kansas. Kansas Herpetological Society Newsletter (120):8-11
2000 Taggart, Travis W. Biogeographic analysis of the reptiles (Squamata) in Ellis County, Kansas. Kansas Herpetological Society Newsletter (121):7-16
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2001 Taggart, Travis W. The KHS 2001 spring field trip: A rainy rendezvous. Kansas Herpetological Society Newsletter (124):12-14
2001 Schmidt, Curtis J. The amphibians, turtles, and reptiles of the Smoky Valley Ranch, Logan County, Kansas. Kansas Herpetological Society Newsletter (124):9-11
2001 Taggart, Travis W. Results of the KHS spring field trip west. Kansas Herpetological Society Newsletter (125):10
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2002 Riedle, J. Daren and A. Hynek. Amphibian and reptile inventory of the Kansas Army Ammunition Plant, Labette County, Kansas. Journal of Kansas Herpetology (2):18-20
2002 Ellis, Mark R. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5
2002 Taggart, Travis W. Results of the spring 2002 KHS field trip. Journal of Kansas Herpetology (3):6-7
To the Cimarron National Grassland, in Morton and Stevens counties.
2002 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (4):15
2002 Rundquist, Eric M. Natural history of the Night Snake, Hypsiglena torquata, in Kansas. Journal of Kansas Herpetology (4):16-20
2002 Reeder, Tod W., Charles J. Cole, and Herbert C. Dessauer Phylogenetic relationships of Whiptail lizards of the genus Cnemidophorus (Squamata: Teiidae): A test of monophyly, reevaluation of karyotypic evolution, and review of hybrid origins. American Museum Novitates (3365):1-61
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Taggart, Travis W. KHS conducts first systematic road survey. Journal of Kansas Herpetology (6):11-12
2003 Platt, Dwight R. Lizards and snakes (Order Squamata) of Harvey County, Kansas. Journal of Kansas Herpetology (6):13-20
2003 Taggart, Travis W. Results of the 2003 KHS spring field trip to Wilson County. Journal of Kansas Herpetology (6):2-5
2003 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (7):10
2003 Volkmann, Al. Cowley County herp count 1. Journal of Kansas Herpetology (7):7
2003 Taggart, Travis W. Logan County herp count. Journal of Kansas Herpetology (7):8
2003 Suleiman, Gibran. Fort Riley herp count. Journal of Kansas Herpetology (7):9
2004 Schmidt, Curtis J. Natural history and status of the exploited Prairie Rattlesnake (Crotalus viridis) in western Kansas and a herpetofaunal inventory of the Smoky Valley Ranch, Logan County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. pp.
2004 Delisle, Jennifer M. and William H. Busby Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Taggart, Travis W. Kansas Herpetological Society 2004 spring field trip. Journal of Kansas Herpetology (9):2
2004 Taggart, Travis W. Results of the 2004 KHS spring field trip to Logan County. Journal of Kansas Herpetology (10):2-7
2004 Volkmann, Al. Cowley County herp count. Journal of Kansas Herpetology (11):10
2004 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (11):11-12
2004 Taggart, Travis W. Results of the KHS 2004 fall feld trip . Journal of Kansas Herpetology (12):15-16
2005 Taggart, Travis W. Results of the KHS 2005 fall field trip [to Crawford County]. Journal of Kansas Herpetology (16):19-21
2006 Taggart, Travis W. Addendum report to biological inventory of the sandsage prairie near Holcomb, Kansas. Sunflower Electric Cooperative, Hays, Kansas. 31pp.
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2006 Wilgers, Dustin J., Eva A. Horne, Brett K. Sandercock, and Allan W. Volkmann. Effects of rangeland management on community dynamics of the herpetofauna of the tallgrass prairie. 62():378-388
2007 Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5pp.
2007 Taggart, Travis W. A biological inventory of the Sunflower Electric Site near Holcomb, Kansas. Journal of Kansas Herpetology 23():11-16
2008 Taggart, Travis W. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3
2010 Miller, Larry L. Investigation of the Checkered Garter Snake in Kansas with notes on other Amphibians, Reptiles, and Turtles encountered. Kansas Department of Wildlife and Parks, Pratt, Kansas. 31pp.
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Hamilton, Bryan Tyler, Rachel Hart , and Jack W. Sites Jr. Feeding Ecology of the Milksnake (Lampropeltis triangulum, Colubridae) in the Western United States. Journal of Herpetology 46(4):515-522
2013 Miller, Larry L. Wellington Lake Herpetological Survey. Collinsorum 2(1/2):12
2013 Baldwin, Mary Kate. “Herps in Havensville” Herp Surveys. Collinsorum 2(3/4):10
2013 Taggart, Travis W. KHS 2012 Spring Field Trip to Bourbon County State Lake. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2012 Summer Field Trip to Meade County State Park. Collinsorum 2(3/4):3
2013 Taggart, Travis W. KHS 2013 Spring Field Trip to Schermerhorn Park, Cherokee County. Collinsorum 2(3/4):4
2013 Taggart, Travis W. KHS 2013 Summer Field Trip to Coldwater Lake, Comanche County. Collinsorum 2(3/4):5
2013 Taggart, Travis W. KHS 2013 Fall Field Trip to Butler County State Lake. Collinsorum 2(3/4):6
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2014 Taggart, Travis W. Results of the 2014 KHS Fall Field Trip to Woodson County. Collinsorum 3(2-4):12
2014 Taggart, Travis W. Results of the 2014 KHS summer field trip to Morton County and adjacent Colorado, New Mexico, and Oklahoma. Collinsorum 3(2-4):12
2014 Goldberg, Stephen R. Reproduction of Six-lined Racerunners, Aspidoscelis sexlineata (Squamata: Teiidae), from New Mexico. Bulletin of the Chicago Herpetological Society 49(1):8-10
2015 Brown, Kasandra A. Occupancy Modeling Of Herpetofauna And Grassland Nesting Birds At Quivira National Wildlife Refuge. Thesis. Fort Hays State University, Hays, Kansas. 72pp.
2015 Mike Houck. 2015 Fort Riley Herpetofaunal Count final report. Collinsorum 4(1):10-11
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2015 Taggart, Travis W. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3
2015 Taggart, Travis W. Fall Field Trip Held In Washington County. Collinsorum 4(3):4
2016 Pittman, Galen L., Henry S. Fitch, and W. Dean Kettle Vertebrate animals on the Fitch Natural History Reservation (1948-2002) Kansas Biological Survey Report Number 188, Lawrence. 48pp.
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3
2016 Taggart, Travis W. Results of the KHS Summer field trip to Caney River, Chautauqua County, Kansas. Collinsorum 5(2-3):4-5
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2016 Tucker, Derek B., Guarino R. Colli, L. G. Giugliano, S. Blair. Hedges, C. R. Hendry, Emily M. Lemmon, Alan R. Lemmon, Jack W. Sites, Jr., and R. Alex Pyron Tucker, D. B., G. R. Colli, L. G. Giugliano, S. B. Hedges, C. R. Hendry, E. M. Lemmon, A. R. Lemmon, J. W. Sites, Jr., and R. A. Pyron. 2016. Methodological congruence in phylogenomic analyses with morphological support for teiid lizards (Sauria: Teiidae). Mol. Phylogenet. Evol. 103: Methodological congruence in phylogenomic analyses with morphological support for teiid lizards (Sauria: Teiidae) Molecular Phylogenetics and Evolution 103():75–84
2017 Taggart, Travis W. and J. Daren Riedle. A Pocket Guide to Kansas Amphibians, Turtles and Lizards. Great Plains Nature Center, Wichita, Kansas. 69pp.
2017 Mardis, Dexter R. Results from three Herpetofaunal tallies at Wichita State University’s Youngmeyer Ranch in Northwestern Elk County. Collinsorum 6(1):8-10
2017 Taggart, Travis W. Results of the 2017 KHS Spring Field Trip to Elk County, Kansas. Collinsorum 6(2-3):6-8
2017 Schmidt, Curtis J. Herp Count: Southeast Ellis County. Collinsorum 6(2-3):9
2018 Houck, Mike. Herp Count: Fort Riley Military Installation Collinsorum 7(1):17
2020 Daniel, Richard E. and Brian S. Edmond. Atlas of Missouri Amphibians and Reptiles for 2019. Privately printed, Columbia, Missouri. 86pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2020 Livo, Lauren J., Todd L. Wilcox, and Daniel J. Martin. Natural history notes: Aspidoscelis sexlineatus viridis: Hatchling phenology. Herpetological Review 51(2):327
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Neosho County: KHS-2020-01 Collinsorum 9(3):11
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Cherokee County: KHS-2020-02 Collinsorum 9(3):11-12
2021 Riedle, J. Daren. Herp Count: Pratt County: KHS-2020-07 Collinsorum 9(3):12
2021 Schmidt, Curtis J and Avery Schmidt. Herp Count: KHS-2020-4 Collinsorum 9(3):12
2021 Mardis, Dexter R. Herp Count: Sumner County: KHS-2020-08. Collinsorum 9(3):12-13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Hodgeman County: KHS-2020-09. Collinsorum 9(3):13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Seward County: KHS-2020-13. Collinsorum 9(3):13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Stanton County: KHS-2020-10. Collinsorum 9(3):13
2021 Taggart, Travis W and Sarah L Taggart. Herp Count: Comanche County: KHS-2020-15. Collinsorum 9(3):13-14
2021 Schmidt, Curtis J. Herp Count: Russell County: KHS-2020-16. Collinsorum 9(3):14
2021 Schmidt, Curtis J. Herp Count: Russell County: KHS-2020-19. Collinsorum 9(3):14
2021 Taggart, Travis W. Herp Count: Russell County: KHS-2020-24. Collinsorum 9(3):14-15
2021 Taggart, Megan M. and Travis W. Taggart. Herp Count: Seward County: KHS-2020-27. Collinsorum 9(3):15
2021 Taggart, Travis W., Dan Fogell, and Christopher Visser. Herp Count: Russell County: KHS-2020-25. Collinsorum 9(3):15
2021 Schmidt, Curtis J. Herp Count: Ellis County: KHS-2020-32. Collinsorum 9(3):16
2021 Walker, James M., Brian K. Sullivan, and James E. Cordes. Sanctioned nomenclature for triploid parthenogenetic Colorado Checkered Whiptail and for other taxa of the lizard genus Aspidoscelis (Family Teiidae). Herpetological Review 52(3):550–558
Followed Tucker et al. (2016) in recognizing Aspidoscelis as masculine (not feminine as Reeder et al. (2002) asserted) which necessitates the specific name be sexlineatus and not sexlineata.
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