An adult Spotted Chorus Frog from Sumner County, Kansas (iNat: 97981052). Image by iNat user Sarah Smith (smolsadpotato).
A tadpole of Pseudacris clarkii. Image © Altig et al. (2006).
An adult Spotted Chorus Frog from Sedgwick County, Kansas (iNat: 94990681). Image by iNat user (suntesha).
An adult Spotted Chorus Frog from Sumner County, Kansas (iNat: 94990681). Image by Sarah Smith.
AMPHIBIA (Amphibians) ANURA (Frogs) HYLIDAE (Treefrogs and Allies)

Spotted Chorus Frog
Pseudacris clarkii (Baird 1854)
sū-dā'-krĭs — clär-kī


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None
Diagnosis:
The Spotted Chorus Frog is characterized by moist skin, a round snout, a light line along the upper lips, and a pattern of dark-edged pale spots (sometimes merging into stripes) scattered irregularly on the back and limbs. The head, body, and limbs are gray. The spots on the back and limbs are pale green and edged with dark gray or black. Some specimens may have a triangular mark between the eyes the same color as the back spots. A greenish stripe passes through the eyes on each side of the head. The belly is white. Males differ from females by their dark throats.
Adults normally 19-28 mm (¾ to 1­1⁄8 inches) in snout-vent length. The largest Kansas specimen is a male (KU 4515) from Rush County with a snout-vent length of 31 mm (1¼ inches) collected by Theodore E. White on 12 August 1927. This is the maximum length throughout the range (Powell et al., 2016).

Distribution:
South-central portions of the state. Generally confined to the drainages south of, and including the Arkansas and Verdigris rivers.
The observations from Allen, Coffey, Labette, and McPherson counties were reported via the Kansas Anuran Monitoring Program (KAMP). KAMP records are by call only (visual identification is not required). Spotted Chorus Frog calls are easily confused with those of the Boreal Chorus Frogs, and therefore, these records need to be corroborated with specimens.
(,   Museum Voucher) (,   Observation) (,   Literature Record) (,   iNat Record), (  Fossil)
Open icons are questionable records; Click on a marker to view details.
Full range depicted by light shaded red area. Export Google Earth (.kml)
  • Occurrence Summary:  
  • 460
    Records 
  • 302
    Museum Vouchers 
  • 158
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Allen (1); Barber (46); Barton (1); Butler (10); Chautauqua (21); Clark (10); Coffey (2); Comanche (47); Cowley (19); Ellis (16); Finney (3); Ford (3); Gray (13); Greenwood (14); Harper (39); Harvey (1); Haskell (6); Hodgeman (1); Kiowa (12); Labette (12); McPherson (2); Meade (21); Montgomery (11); Ness (1); Pawnee (12); Pratt (3); Rush (7); Russell (1); Sedgwick (39); Seward (2); Sumner (82); Trego (1); Unknown (1);

Fossil History:
Reported from the WaKeeney Local Fauna of Trego County based on a single right ilium (UMMP V55414) missing the anterior portion of the shaft by Wilson (1968) and an ilium (MSU-VP 764) by Holman (1976). The WaKeeney Local Fauna site is considered Lower Pliocene: Middle or Late Clarendonian (13,600,000 to 10,300,000 years BP). This site is marginally extralimitial of the species current range, however Chantell (1964) reported a fossil specimen from the Norden Bridge Local Fauna (Upper Miocene: Valentine Formation) in Brown County, Nebraska.

Natural History:
They are found in open prairie grasslands and pastures. They breed in shallow ponds, cattle tanks, and ephemeral streams. As winter approaches, they burrow under thatch and loose soil until March and April rains bring them out again. The call is a raspy trill wrraaay-wrraaay-wrraaay repeated 20-30 times rapidly. They call from March into June.
The Spotted Chorus Frog inhabits open prairie grasslands and the edges of woodlands. It is most active during spring and summer rains and becomes very secretive during dry weather, probably retreating to underground burrows. This small frog is active from March to September. Little is known of its habits in Kansas.
The Spotted Chorus Frog breeds opportunistically from March to September, depending on rainfall and suitable temperatures. Apparently, it breeds in open, shallow, temporary rain pools, although mating sometimes occurs in permanent ponds. Males chorus and attract females to breed. Clasping may occur for an entire day, and each female deposits up to 1,000 eggs. The eggs are laid in small masses of 6- 30 each on grasses and stems just below the water surface. The eggs hatch in two to ten days, and the tadpoles remain in the pond until they metamorphose in late summer.
Fleharty and Johnson (1974) reported the Spotted Chorus Frog in a breeding chorus together with American Toads and Leopard Frogs during mid-April in Chautauqua County. The frogs were calling from an alfalfa field flooded by recent rains. Gray and Stegall (1979) found a single example of this frog in a cave in the Red Hills on 29 September. I observed a breeding chorus of these frogs, together with choruses of Plains Spadefoots and Great Plains Toads, by day in a roadside ditch in Comanche County on 11 May.
No specific information on the food preferences of this frog in Kansas is available, but it probably eats small insects.
Predators of the Spotted Chorus Frog include larger frogs, large birds, and snakes.

Occurrence Activity:
Number of Unique Obervations (=days): 103; Range: 17 Mar to 13 Oct
Chorusing:

Audio recording by Keith Coleman.

Chorusing Phenology: The black outlined dots denote the Julian date (day of the year; 1 January = 1 to 31 December = 365) an observation was made. The thin red line depicts the range of dates between the beginning of the first, and end of the fourth quartile (excluding outliers; Tukey method). The thick light blue bar represents the second and third quartile (interquartile range; the middle 50% of all observations). Only one observation per Julian date is included in the graphs; so a date with multiple observations carries the same weight as a date with only one observation. The vertical bars correspond to the 12 months of the year; January through December.
Number of Unique Obervations: 163; Range: 22 Mar to 28 Aug; Interquartile range: 22 Apr to 03 Jun;

Remarks:
The Spotted Chorus Frog was first reported in Kansas by Smith, (1933) in his Master's thesis as Pseudacris nigrita. The earliest existing specimen (USNM 45828) was collected at Kiowa (Barber County) by famed Bureau of Biological Survey naturalist Vernon O. Bailey on 31 August 1892.
The records mapped for Ellis and Russell counties by Collins (1993) is in need of verification. There are pockets of Boreal Chorus Frogs along Big Creek and the Smoky Hill River in these two counties which possess broken striping and resemble Spotted Chorus Frog. These northern spotted Pseudacris do not have green blotches as do most Spotted Chorus Frogs, and instead, are colored as typical Boreal Chorus Frogs. Live specimens are needed to verify their presence.Smith (1933) reported Pseudacris nigrita from Kansas based on six specimens (KU 4511-6) from Rush County. By the time his thesis was published a year later (Smith, 1934) he considered these specimens to be P. clarkii, though he remarked on their confounding characters and placement, which is still in debate today (Moriarty and Cannatella, 2004).
Moriarty and Cannatella (2004), Moriarty (2007), and Moriarty et al. (2008) found that Pseudacris clarkii and Pseudacris maculata were not reciprocally monophyletic.

Bibliography:
1922 O'Roke, Earl C. Frogs and frogging. Transactions of the Kansas Academy of Science 30():448-451
1933 Smith, Hobart M. The Amphibians of Kansas. Thesis. University of Kansas, Lawrence. 383pp.
The first full accounting of the twenty-five species of amphibians known to occur in Kansas. Includes Ambystoma maculatum which is currently not included in the Kansas faunal list.
1934 Smith, Hobart M. The Amphibians of Kansas. American Midland Naturalist 15(4):377-527
The formal publication of Hobart Smith's Master's Thesis (Smith 1933), though there are several updated and additions. In addition to the species accounts for all twenty-five species, the paper includes a history of amphibian biology in Kansas and discussions on taxonomy and physiography. 
1936 Brumwell, Malcolm J. Distributional records of the reptilia and amphibians of Kansas. Privately printed, . 22pp.
County dot maps of the Kansas herpetofauna. This work has been attributed to have been written around 1933, but that may be in error. 
Hypsiglena jani was not known from Kansas until Claude W. Hibbard collected three specimens on the Stevenson Ranch in north-central Clark County (above Clark State Lake) during June 1936 (Hibbard, 1937). Brumwell plotted this locality, which leads me to believe that the 1936 would have been the earliest date this manuscript could have been written.
1948 Eaton, Theodore H., Jr., and R. M. Imagawa. Early development of Pseudacris clarkii. Copeia 1948():263-266
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
The first modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table and text say 97 on p. 10) and 13 "probable but unverified" species and subspecies.
1953 Bellis, Edward D. The effects of temperature on the breeding calls of some Oklahoma Salientians. Thesis. University of Oklahoma, Norman. pp.
1953 Schmidt, Karl P. A Check List of North American Amphibians and Reptiles. 6th Edition. University of Chicago Press, Chicago, Illinois. 280pp.
Schmidt's first edition of his standardized checklist to North American amphibians and reptiles. Includes several specific references to Kansas in the range descriptions.
1956 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. Second edition. University of Kansas Museum of Natural History Miscellaneous Publication (9):1-356
Hobart M. Smith's updated second edition of his first (1950) modern herpetology of Kansas. Includes locality dot maps within individual species accounts. Reports 96 species from Kansas (table says 97 on p. 10; text says 98 on p. 10) and 11 "probable but unverified" species and subspecies. The second edition has updated taxonomy, added Plestiodon laticeps, and removed Eurycea tynerensis.
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
Examined 2,628 Kansas reptiles of 48 species consisting of 27 turtles of 4 species, 1,736 lizards of 12 species and 892 snakes of 32 speciesfor chiggers. Eleven species of chiggers were recovered from reptiles.
For amphibians, 1188 individuals of 21 species were examined. Five species of chigger mite were recovered from amphibians.
1957 Bellis, Edward D. The effects of temperature on salientian breeding calls. Copeia 1957(2):85-89
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1966 Chantell, Charles J. Late Coenozoic hylids from the Great Plains. Herpetologica 22(4):259-264
1967 Gier, Herschel T. Vertebrates of the Flint Hills. Transactions of the Kansas Academy of Science 70(1):51-59
1968 Wilson, Richard L. Systematics and faunal analysis of a Lower Pliocene vertebrate assemblage from Trego County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 22():75-126
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
Joseph T. Collins first Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Smith 1956)>
1974 Fleharty, Eugene D. and Jerry D. Johnson. Distributional records of herptiles from the Chautauqua Hills of southeastern Kansas. Transactions of the Kansas Academy of Science 77(1):65-67
1975 Rundquist, Eric M. Amphibians and Reptiles of Kingman County, Kansas. Privately Printed, Lawrence, Kansas. 3pp.
Short accounts for twenty-nine recognized amphibians and reptiles from Kingman County, Kansas. With habitat descriptions and for some species, estimates of population density.
1975 Holman, J. Alan. Herpetofauna of the WaKeeney local fauna (Lower Pliocene: Clarendionian) of Trego County, Kansas. Pages 49-66 in Studies on Cenozoic Paleontology and Stratigraphy in honor of Claude W. Hibbard. Museum of Paleontology, The University of Michigan, Ann Arbor, Michigan. pp.
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1977 Perry, Janice. Kansas herps needed. Kansas Herpetological Society Newsletter (18):2-3
List of Kansas amphibians and reptiles desired for the SSAR/HL meeting to be held 7-13 August 1977.
1977 Grow, David. Clark County visited by the Society. Kansas Herpetological Society Newsletter (19):1-2
1978 Warner, M. and R. Wencel. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16
1978 Capron, Marty B. Four county collecting raid: A south central Kansas herping saga. Kansas Herpetological Society Newsletter (26):9-12
1978 Collins, Joseph T. and Janalee P. Caldwell. New records of fishes, amphibians, and reptiles in Kansas for 1977. Technical Publication of the State Biological Survery of Kansas 6():70-88
1979 Gray, Peter and Eddie Stegall. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
A summary of known information on the amphibian, reptile, and fish faunas of the Arkansas River above Great Bend. The report details associated field activities for procuring fish, however no new surveys for amphibians and reptiles were undertaken. Information on herps from Finney County was provided by Michael Rush (FHSU) and thus made available before the publication of his thesis (Rush, 1981). The western Arkansas River drainage had experienced little attention by herpetologists before this study, and the species accounts reflect that paucity of data. Additionally, the report omits several older records (e.g. for Anaxyrus debilisThamnophis cyrtopsis, and Lampropeltis calligaster) from the westernmost reaches of the Arkansas River drainage in Kansas.
1980 Corn, Paul S. Comment on the occurrence of Pseudacris clarki in Montana. Bulletin of Chicago Herpetological Society 15():77-78
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
Joseph T. Collins second Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1974)>
1983 Trott, Gene. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (52):3-4
1984 Secor, Stephen M. and Charles C. Carpenter. Distribution maps of Oklahoma reptiles. Oklahoma Herpetological Society Special Publication (3):1-57
1986 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1985. Kansas Herpetological Society Newsletter (63):4
1987 Simmons, John E. September 1987 field trip report. Kansas Herpetological Society Newsletter (69):42894
1988 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1987. Kansas Herpetological Society Newsletter (71):13-19
1988 Miller, Larry L. Harper County KHS field trip well attended. Kansas Herpetological Society Newsletter (72):5-6
1989 Collins, Joseph T. First Kansas herp counts held in 1989. Kansas Herpetological Society Newsletter (77):11-
1990 Lardie, Richard L. Kansas threatened species and protection of the Gypsum Hills habitat. Kansas Herpetological Society Newsletter (80):14-15
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1990 Pierce, Benjamin A. and Patricia H. Whitehurst. Pseudacris clarkii. Catalogue of American Amphibians and Reptiles (458):1-3
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1992 Ford, Kenneth M. III. Herpetofauna of the Albert Ahrens Local Fauna (Pleistocene: Irvingtonian), Nebraska. Thesis. Michigan State University, Lansing, Michigan. 44pp.
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Gubanyi, James E. An observation on the stomach contents of a Texas Longnose Snake (Rhinocheilus lecontei tessellatus). Kansas Herpetological Society Newsletter (89):17
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
Joseph T. Collins third Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1982)>
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
See, 1994 Rundquist, Eric M. Additions and corrections [to the results of the sixth annual KHS herp counts held 1 April-31 May 1994]. Kansas Herpetological Society Newsletter (98):4.
1995 Moriarty, Emily C. and Joseph T. Collins. First known occurrence of amphibian species in Kansas. Kansas Herpetological Society Newsletter (100):28-30
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1996 Young, Eugene A. and Max C. Thompson. Waterbird usage of the Warner Marsh, Slate Creek Wetlands, Sumner County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 80pp.
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1996 Shoup, J. Mark. Treefrogs, indeed! Kansas Wildlife and Parks 53(4):34
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1997 Rundquist, Eric M. Addendum to 1997 KHS herp counts. Kansas Herpetological Society Newsletter (109):14-15
1997 da Silva, Hlio Ricardo. Two character states new for Hylines and the taxonomy of the genus Pseudacris. Journal of Herpetology 31(4):609-613
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. A Key to Amphibians & Reptiles of the Continental United States and Canada. Univ Press of Kansas, Lawrence, Kansas. 131pp.
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2003 Taggart, Travis W. Kansas Herpetological Society 2003 spring field trip. Journal of Kansas Herpetology (5):3-4
2003 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (7):10
2004 Delisle, Jennifer M. and William H. Busby. Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (11):11-12
2004 Moriarty, Emily C. and David C. Cannatella. Phylogenetic relationships of the North American Chorus Frogs (Pseudacris: Hylidae) Molecular Phylogenetics and Evolution 30():409-420
2005 Lannoo, Michael (Editor) Amphibian Declines: The Conservation Status of United States Species. University fo California Press, Berkeley. 1115pp.
2006 Altig, Ronald, Roy W. McDiarmid, Kimberly A. Nichols, and Paul C. Ustach. Tadpoles of the United States and Canada: A Tutorial and Key Electronic files accessible at http://www.pwrc.usgs.gov/tadpole/. Patuxent Wildlife Research Center, Laurel, MD, USA. ():
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2007 Moriarty-Lemmon, Emily. Patterns and Processes of Speciation in North American Chorus Frogs (Pseudacris). Dissertation. University of Texas, Austin. 304pp.
2008 Taggart, Travis W. KHS 2008 spring field trip. Journal of Kansas Herpetology (25):2-3
2008 Moriarty-Lemmon, Emily, Alan R. Lemmon, Joseph T. Collins, and David C. Cannatella. A new North American chorus frog species (Amphibia: Hylidae: Pseudacris) from the south-central United States. Zootaxa (1675):1-30
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
Joseph T. Collins fourth Kansas herpetology. <Need to get species total and principal differences with previous 'version' (= Collins 1993)>
2010 Murrow, Daniel G. Kansas Herpetological Society spring field trip. Journal of Kansas Herpetology (33):2-3
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to be held in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2012 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada: Second Edition, Revised and Updated. Univ Press of Kansas, Lawrence, Kansas. 152pp.
2013 Dodd, C. Kenneth. Frogs of the United States and Canada. John Hopkins University Press, Baltimore, Maryland. 982pp.
2013 Miller, Larry L. Wellington Lake Herpetological Survey. Collinsorum 2(1/2):12
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2014 Barrow, Lisa N., Hannah F. Ralicki, Sandra A. Emme, Emily Moriarty Lemmon. Species tree estimation of North American chorus frogs (Hylidae: Pseudacris) with parallel tagged amplicon sequencing. Molecular Phylogenetics and Evolution 75():78-90
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2017 Taggart, Travis W. and J. Daren Riedle. A Pocket Guide to Kansas Amphibians, Turtles and Lizards. Great Plains Nature Center, Wichita, Kansas. 69pp.
2017 Crother, Brian I. (editor) Scientific and Standard English Names of Amphibians and Reptiles of North America North of Mexico, with Comments Regarding Confidence in Our Understanding. Eighth edition. Herpetological Circulars (43):1-102
2019 Powell, Robert, Joseph T Collins, and Errol D Hooper Jr. Key to the Herpetofauna of the Continental United States and Canada. Third Edition. Univ Press of Kansas, Lawrence, Kansas. 192pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
2020 Goldberg, Stephen R. Notes on reproduction of the Spotted Chorus Frog, Pseudacris clarkii (Anura: Hylidae), from Oklahoma and Texas. Bulletin of the Chicago Herpetological Society 55(10):197-198
2020 Banker, Sarah E., Alan R. Lemmon, Alyssa Bigelow Hassinger, Mysia Dye, Sean D. Holland, Michelle L. Kortyna, Oscar E. Ospina, Hannah Ralicki, and Emily Moriarty Lemmon. Hierarchical hybrid enrichment: Multitiered genomic data collection across evolutionary scales, with application to Chorus Frogs (Pseudacris). Systematic Biology 69(4):756-773
2021 Ethier, Jeffery P., Aurore Fayard, Peter Soroye, Daeun Choi, Marc. J. Mazerolle, and Vance L. Trudeau. Life history traits and reproductive ecology of North American chorus frogs of the genus Pseudacris (Hylidae). Frontiers in Zoology 18(40):18
2023 Dodd, C. Kenneth. Frogs of the United States and Canada. Second Edition. John Hopkins University Press, Baltimore, Maryland. 1032pp.
2023 Russell, Elisabeth Habitat associations and fine-scale movements of the Red-spotted Toad (Anaxyrus punctatus) in Kansas and the efficacy of remote telemetry for monitoring small-scale movements. Thesis. Fort Hays State University, Hays, Kansas. 81pp.
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Travis W. Taggart © 1999-2025 — w/ Sternberg Museum of Natural History, Fort Hays State University