GREAT PLAINS TOAD
Anaxyrus cognatus (Say 1822: 190)
ăn-ăx-ī'-rŭs — käg-nā'-tŭs


Conservation Status:

State: None

Federal: None
NatureServe State: S5 - Secure
NatureServe National: N5 - Secure
NatureServe Global: G5 - Secure
CITES: None

An adult female Great Plains Toad from Hamilton County, Kansas. © Travis W. Taggart.
An adult specimen from Clark County. Image © Suzanne L. Collins, CNAH.
A tadpole of Bufo cognatus. Image © Altig et al. (2006).
A chorusing adult male from Seward County. Image © Travis W. Taggart.

Description:
The Great Plains Toad has large paired dark (black, brown, or green) blotches on its back. Each blotch is boldly bordered by lighter pigment and contains many warts. Each blotch contains several small warts. The colors of this toad are generally yellowish, brown, greenish, or gray on top. Below is unspotted, cream to white, with a yellow or orange-yellow seat patch. Some specimens have been found with a narrow, light mid-dorsal stripe. The head of the Great Plains toad is relatively small with a well-developed cranial crest. Their snout is blunt and rounded.
Adults normally 50- 85 mm (2 - 3 3/8 inches) in SVL; largest Kansas specimen: female (KU 186099) from Sumner County with SVL of 102 mm (4 inches) collected by Jeff Ehlers on 23 June 1980; maximum length throughout range: 4½ inches (Conant and Collins, 1998).

Distribution:
Found throughout western Kansas to the east edge of the Flint Hills. But also follows the riparian corridor along the Kansas River and then along the Missouri River to the North. Specimens from southern Douglas and northwest Miami counties are each over 50 years old and in need of verification.
Range-wide 

(, Museum Voucher) (, Observation) (, Literature Record)
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  • Occurrence Summary:  
  • 1,091
    Records 
  • 839
    Museum Vouchers 
  • 252
    Other Observations 
Some county occurrences indicated below may be too imprecise to map above.
County Breakdown: County Name (# occurrences):
Atchison (1); Barber (18); Barton (24); Brown (1); Butler (3); Chase (1); Chautauqua (1); Cheyenne (4); Clark (6); Clay (1); Cloud (1); Comanche (20); Cowley (68); Dickinson (2); Doniphan (1); Douglas (71); Edwards (3); Elk (2); Ellis (234); Ellsworth (2); Finney (31); Ford (8); Geary (5); Gove (4); Graham (43); Grant (5); Gray (8); Greeley (5); Hamilton (2); Harper (13); Harvey (4); Haskell (1); Hodgeman (3); Jefferson (9); Jewell (1); Johnson (2); Kearney (9); Kingman (15); Kiowa (13); Lane (3); Leavenworth (3); Logan (2); Marion (4); Marshall (4); McPherson (10); Meade (55); Miami (1); Mitchell (1); Morton (8); Ness (53); Norton (8); Ottawa (1); Pawnee (21); Phillips (8); Pottawatomie (3); Pratt (4); Rawlins (2); Reno (27); Republic (1); Rice (12); Riley (4); Rooks (2); Rush (15); Russell (1); Saline (3); Scott (23); Sedgwick (18); Seward (25); Shawnee (5); Sheridan (1); Sherman (2); Stafford (42); Stanton (1); Stevens (4); Sumner (20); Thomas (9); Trego (22); Unknown (4); Wabaunsee (1); Wallace (6); Washington (3); Wichita (2); Wyandotte (2);

Natural History:
Following spring rains from April to June, male Great Plains Toads call from semi-permanent ponds to attract mates. The males use their frontward-projecting, sausage-shaped vocal sac to produce their call, which is a loud, metallic trill that lasts 20-50 seconds. A female may lay up to 20,000 eggs, which hatch in two to three days. Eggs are laid in strings. The young toads transform and leave the water four to six weeks later.
Hartman (1906) reported a diet of snout-beetles and dung-beetles in Kansas specimens, though they likely will consume any live small animal that fits in their mouth.


Occurrence Activity:
The blue dates denote chorusing actity. The red dates are other occurrences. The darker a date is, the greater the relative number of observations for that date.
Chorusing:

Audio recording by Keith Coleman.

Chorusing Phenology: The black outlined dots denote the Julian date (day of the year; 1 January = 1 to 31 December = 365) an observation was made. The thin red line depicts the range of dates between the beginning of the first, and end of the fourth quartile (excluding outliers; Tukey method). The thick light blue bar represents the second and third quartile (interquartile range; the middle 50% of all observations). Only one observation per Julian date is included in the graphs; so a date with multiple observations carries the same weight as a date with only one observation. The vertical bars correspond to the 12 months of the year; January through December.
# Unique Obervations: 212; Range: 27 Mar to 31 Aug; Interquartile range: 11 May to 24 Jun;

Observation Type: (of recorded types)
Remarks:
First reported in Kansas by Cragin (1880). The earliest extant specimen from Kansas is MCZ 20141 (1881). Smith (1946) described the tadpoles of this species from a series taken at Meade County State Park.
Gloyd (1929) reported a large female from Riley County that was collected at approximately 10 pm on a small road near the Blue River NE of Manhattan on 12 July 1928.
Fossils -
Miocene (Clarendonian NALMA): WaKeeney Local Fauna, Trego County (Wilson, 1968; Holman, 1975; Sanchiz, 1998);
Miocene (Hemphillian NALMA): Quarry E, near Long Island, Phillips County (Tihen, 1962, Gehlbach, 1965), Sanchiz (1998);
Pliocene (Blancan NALMA): Rexroad Local Fauna, Meade County (Tihen, 1962, Gehlbach, 1965, Sanchiz, 1998); White Rock Fauna, Republic County (Eshelman, 1975, Sanchiz, 1998);
Pleistocene (Rancholabrean NALMA): Butler Spring Local Fauna, Meade County (Tihen 1962, Gehlbach 1965, Holman 1986, Holman 1995, Sanchiz 1998); Cragin Quarry Local Fauna, Meade County (Tihen 1962, Holman 1995, Sanchiz 1998; Jinglebob Local Fauna, Meade County (Tihen 1954, Tihen 1962, Holman 1995, Sanchiz 1998; Jones Local Fauna, Meade County (Tihen 1962, Gehlbach 1965, Holman 1995, Sanchiz 1998); Sandahl Local Fauna, McPherson County (Holman 1971, Holman 1995, Sanchiz 1998).


Bibliography:
1822 Say, Thomas. In James, Edwin. Pages 1783 in Account of an expedition from Pittsburgh to the Rocky Mountains, performed in the years 1819, '20, by order of the Hon. J. C. Calhoun, Secretary of War, under the command of Major Stephen H. Long. Vol. 2. H. C. Carey & I. Lea., Philadelphia. pp.
1880 Cragin, Francis W. A preliminary catalogue of Kansas reptiles and batrachians Transactions of the Kansas Academy of Science 7():114-123
1882 Yarrow, Henry C. Check list of North American Reptilia and Batrachia with catalogue of specimens in U. S. National Museum. Bulletin of the United States National Museum (24):1-249
1887 Garman, Samuel. Reptiles and batrachians from Texas and Mexico. Bulletin of the Essex Institute 19():119-138
1906 Dickerson, Mary C. The Frog Book; North American Toads and Frogs, with Study of the Habits and Life Histories of Those of the Northeastern States. Doubleday, Page & Company, New York, New York. 253pp.
1906 Hartman, Frank A. Food habits of Kansas lizards and batrachians. Transactions of the Kansas Academy of Science 20():225-229
1925 Storer, Tracy I. A synopsis of the amphibia of California. University of California Publications in Zoology (27):1-342
1929 Gloyd, Howard K. Two additions to the herpetological fauna of Riley County, Kansas. Science 66(1776):44
1929 Taylor, Edward H. List of reptiles and batrachians of Morton County, Kansas, reporting species new to the state fauna. University of Kansas Science Bulletin 19(6):63-65
1933 Smith, Hobart M. The Amphibians of Kansas Thesis. University of Kansas, Lawrence. 383pp.
1933 Stejneger, Leonhard and Thomas Barbour. A Checklist of North American Amphibians and Reptiles. 3rd Edition. Harvard University Press, Cambridge, Massachusetts. pp.
1934 Brennan, Lawrence A. A check list of the amphibians and reptiles of Ellis County, Kansas. Transactions of the Kansas Academy of Science 37():189-191
1935 Brennan, Lawrence A. Notes on the Distribution of Amphibia and Reptilia of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 114pp.
1937 Brennan, Lawrence A. A study of the habitat of reptiles and amphibians of Ellis County, Kansas. Transactions of the Kansas Academy of Science 40():341-347
1942 Hudson, G. E. The amphibians and reptiles of Nebraska. Nebraska Conservation Bulletin 24():1-146
1944 Marr, John C. Notes on amphibians and reptiles from the central United States. American Midland Naturalist 32(2):478-490
1946 Smith, Hobart M. The tadpoles of Bufo cognatus Say. University of Kansas Publications Museum of Natural History 1(3):93-96
1950 Smith, Hobart M. Handbook of Amphibians and Reptiles of Kansas. University of Kansas, Museum of Natural History, Miscellaneous Publication (2):336
1956 Loomis, Richard B. The chigger mites of Kansas (Acarina, Trombiculidae). University of Kansas Science Bulletin 37():1195-1443
1958 Blair, W. Frank. Mating call in the speciation of anuran amphibians. The American Naturalist 92(862):27-51
1962 Gish, Charles D. The Herpetofauna of Ellis County, Kansas. Thesis. Fort Hays State University, Hays, Kansas. 34pp.
1968 Wilson, Richard L. Systematics and faunal analysis of a Lower Pliocene vertebrate assemblage from Trego County, Kansas. Contributions from the Museum of Paleontology, University of Michigan 22():75-126
1971 Holman, J. Alan. Herpetofauna of the sandahl local fauna (Pleistocene: Illinoian) of Kansas. Contributions from the Museum of Paleontology, University of Michigan 23(22):349-355
1974 Collins, Joseph T. Amphibians and Reptiles in Kansas University of Kansas Museum of Natural History Public Education Series (1):283 pp
1974 Perry, Janice. KHS members take trip to southwest Kansas. Kansas Herpetological Society Newsletter (3):2-3
1975 Holman, J. Alan. Herpetofauna of the WaKeeney local fauna (Lower Pliocene: Clarendionian) of Trego County, Kansas. Papers on Paleontology, University of Michigan 1974(12):49-66
1975 Rundquist, Eric M. First KHS field trip yields three county records. Kansas Herpetological Society Newsletter (7):1-3
1975 Taylor, Edward H., A. Byron Leonard, Hobart M. Smith, and George R. Pisani. Edward H. Taylor: Recollections of an Herpetologist . Monograph of the Museum of Natural History, the University of Kansas (4):1-159
1976 Capron, Marty and Jan Perry. A July weekend in Great Bend. Kansas Herpetological Society Newsletter (14):1-2
1976 Rundquist, Eric M. Field checklist (of) amphibians and reptiles of Kansas. Kansas Herpetological Society, Lawrence. pp.
1977 Irwin, Kelly J. KHS Ottawa County meeting profitable. Kansas Herpetological Society Newsletter (20):1-2
1977 Knight, James L. and Joseph T. Collins. The amphibians and reptiles of Cheyenne County, Kansas, Report Number 15. Kansas Biological Survey, Lawrence. 19pp.
1977 Stegall, Eddie. First Strecker's Chorus Frog collected in Kansas. Kansas Herpetological Society Newsletter (21):11-13
1977 Trott, Gene. Chikaskia River wildlife study. Kansas Herpetological Society Newsletter (19):2-3
1978 Curl, Richard L. Final Environmental Statement: Milford Lake Kansas operation and maintenance. US Army Corps of Engineers, Kansas City District. 158pp.
1978 Warner, M. and R. Wencel. Chikaskia River study held near Caldwell. Kansas Herpetological Society Newsletter (25):15-16
1979 Gray, Peter and Eddie Stegall. A field trip to the Red Hills. Kansas Herpetological Society Newsletter (29):6-8
1979 Miller, Larry L. Life in Kansas and Oklahoma once again threatened by pesticides. Kansas Herpetological Society Newsletter (31):7
1980 Clarke, Robert F. Herptiles and fishes of the western Arkansas River in Kansas. United States Army Corps of Engineers, Albuquerque, New Mexico. 55pp.
1980 Guarisco, Hank. Record size Great Plains Toad collected in Sumner County. Kansas Herpetological Society Newsletter (38):5
1980 Kunz, T. H., J. R. Choate, and S. B. George Distributional records for three species of mammals in Kansas Transactions of the Kansas Academy of Science 83():74-77
1982 Collins, Joseph T. Amphibians and Reptiles in Kansas. 2nd edition. University of Kansas Museum of Natural History Public Education Series (8):
1983 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1982 . Technical Publication of the State Biological Survey of Kansas 13():9-21
1984 Collins, Joseph T. New records of fishes, amphibians, and reptiles in Kansas for 1983. Kansas Herpetological Society Newsletter (56):15-26
1986 Holman, J. Alan. Butler Spring herpetofauna of Kansas (Pleistocene: Illinoian) and its climatic significance. Journal of Herpetology 20():568-569
1988 Nulton, Michael T. and Michael S. Rush. New county records of amphibians and reptiles in Gray County, Kansas. Kansas Herpetological Society Newsletter (74):10-12
1989 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1988. Kansas Herpetological Society Newsletter (75):15-18
1990 Collins, Joseph T. Maximum size records for Kansas amphibians and reptiles. Kansas Herpetological Society Newsletter (81):13-17
1990 Collins, Joseph T. Results of second Kansas herp count held during April-May 1990. Kansas Herpetological Society Newsletter (81):10-12
1990 Krupa, James J. Bufo cognatus. Catalogue of American Amphibians and Reptiles (457):1-8
1991 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1990. Kansas Herpetological Society Newsletter (83):7-13
1991 Collins, Joseph T. Results of third Kansas herp count held during April-May 1991. Kansas Herpetological Society Newsletter (85):9-13
1991 Collins, Joseph T. and Suzanne L. Collins. Reptiles and Amphibians of the Cimarron National Grasslands, Morton County, Kansas. U. S. Forest Service, Elkhart, Kansas. 60pp.
1991 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed. Houghton Mifflin Company, Boston, Massachusetts. pp.
1992 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1991. Kansas Herpetological Society Newsletter (87):12-17
1992 Collins, Joseph T. Results of the fourth Kansas herp count held during April-May 1992. Kansas Herpetological Society Newsletter (89):10-
1992 Taggart, Travis W. Results of the KHS annual field trip to Sheridan County State Lake. Kansas Herpetological Society Newsletter (90):3-4
1993 Collins, Joseph T. and Rundquist, Eric M. Results of the fifth Kansas herp count held during April-June 1993 . Kansas Herpetological Society Newsletter (94):7-11
1993 Collins, Joseph T. and Suzanne L. Collins. Amphibians and Reptiles in Kansas. Third Edition. University Press of Kansas, Lawrence, Lawrence. 397pp.
1994 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1993. Kansas Herpetological Society Newsletter (97):15-19
1994 Rundquist, Eric M. Results of the sixth annual KHS herp counts held 1 April-31 May 1994. Kansas Herpetological Society Newsletter (97):5-14
1995 Anderson, Lewis, Mark Shaw, Jeff Blodig, and Tom Walker. Report to the Kansas Department of Wildlife and Parks: Herps encountered during REmap project, summer 1994. Kansas Herpetological Society Newsletter (99):10-17
1995 Holman, J. Alan. Pleistocene Amphibians and Reptiles. Oxford University Press, New York. 243pp.
1995 Moriarty, Emily C. and Joseph T. Collins. First known occurrence of amphibian species in Kansas. Kansas Herpetological Society Newsletter (100):28-30
1995 Rundquist, Eric M. Additional KHS herp counts for 1995. Kansas Herpetological Society Newsletter (102):11-
1995 Rundquist, Eric M. Results of the seventh annual KHS herp counts held 1 April-31 May 1995. Kansas Herpetological Society Newsletter (101):11-17
1996 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1995. Kansas Herpetological Society Newsletter (103):13-15
1996 Dundee, Harold A. Some reallocations of type localities of reptiles and amphibians described from the Major Stephen H. Long Expedition to the Rocky Mountains, with comments on some of the statements made in the account written by Edwin James.  Tulane Studies in Zoology and Botany 30():75–89
1996 Rakestraw, J. Spring herp counts: A Kansas tradition. Reptile & Amphibian Magazine (March-April):75-80
1996 Rundquist, Eric M. Notes on the natural history of some Kansas amphibians and reptiles: Parasites. Kansas Herpetological Society Newsletter (105):16-17
1996 Rundquist, Eric M. Results of the eighth annual KHS herp counts Held 1 April-31 May 1996. Kansas Herpetological Society Newsletter (104):6-17
1997 Rundquist, Eric M. Addendum to 1997 KHS herp counts. Kansas Herpetological Society Newsletter (109):14-15
1997 Rundquist, Eric M. Results of the ninth annual KHS herp counts held 1 April-31 May 1997. Kansas Herpetological Society Newsletter (108):12-17
1998 Conant, Roger and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. 3rd ed, expanded. Houghton Mifflin Company, Boston, Massachusetts. pp.
1998 Rundquist, Eric M. Results of the tenth annual KHS herp counts for 1998, held 1 April-31 May. Kansas Herpetological Society Newsletter (112):11-18
1999 Rundquist, Eric M. Kansas Herpetological Society herp counts: A 10 year summary and evaluation. Kansas Herpetological Society Newsletter (115):42962
2000 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 1999. Kansas Herpetological Society Newsletter (119):7-9
2000 Rundquist, Eric M. Results of the eleventh and twelfth annual KHS herpetofaunal counts for 1999-2000, held 1 April-31 May. Kansas Herpetological Society Newsletter (122):11-16
2000 Van Doren, Mark D. and Curtis J. Schmidt. A herpetological survey of the Fort Larned National Historic Site, Pawnee County, Kansas. Kansas Herpetological Society Newsletter (120):8-11
2001 Rundquist, Eric M. Results of the thirteenth annual KHS herp counts for 2001, held 1 April-30 June. Kansas Herpetological Society Newsletter (125):13-16
2002 Collins, Joseph T. New records of amphibians and reptiles in Kansas for 2001. Journal of Kansas Herpetology (1):10-11
2002 Ellis, Mark R. Fall 2002 KHS field trip to Washington County. Journal of Kansas Herpetology (2):4-5
2002 Kingsbury, Bruce and Joanna Gibson. Habitat Management Guidelines for Amphibians and Reptiles of the Midwest. Publication of Partners in Amphibian and Reptile Conservation, Address not given. 152pp.
2003 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (7):10
2003 Suleiman, G. Fort Riley herpetofaunal count. Journal of Kansas Herpetology (5):11-12
2003 Taggart, Travis W. Kansas Herpetological Society 2003 spring field trip. Journal of Kansas Herpetology (5):3-4
2003 Taggart, Travis W. Logan County herp count. Journal of Kansas Herpetology (7):8
2004 Delisle, Jennifer M. and William H. Busby Biological inventory for vertebrates at Fort Larned National Historic Site of the southern plains network. Natural Heritage Inventory, Kansas Biological Survey, Lawrence. 61pp.
2004 Miller, Larry L. Sumner County herp count. Journal of Kansas Herpetology (11):11-12
2004 Schmidt, Curtis J. Geographic distribution: Bufo cognatus. Kansas. Journal of Kansas Herpetology (11):13
2004 Taggart, Travis W. Kansas Herpetological Society 2004 spring field trip. Journal of Kansas Herpetology (9):2
2006 Altig, Ronald, Roy W. McDiarmid, Kimberly A. Nichols, and Paul C. Ustach Tadpoles of the United States and Canada: A Tutorial and Key Electronic files accessible at http://www.pwrc.usgs.gov/tadpole/. Patuxent Wildlife Research Center, Laurel, MD, USA. ():
2006 Frost, Darrel R, Taran Grant, Julian Faivovich, Raoul H. Bain, Alexander Haas, Celio F. B. Haddad, Rafael O. De Sa, Alan Channing, Mark Wilkinson, Stephen C. Donnellan, Christopher J. Raxworthy, Jonathan A. Campbell, Boris L. Blotto, Paul Moler, Robert C. Drewes, Ronald A. Nussbaum, John D. Lynch, David M. Green, and Ward C. Wheeler. The amphibian tree of life Bulletin of the American Museum of Natural History (297):370
2006 Taggart, Travis W. Addendum report to biological inventory of the sandsage prairie near Holcomb, Kansas. Sunflower Electric Cooperative, Hays, Kansas. 31pp.
2006 Taggart, Travis W. Distribution and status of Kansas herpetofauna in need of information. State Wildlife Grant T7. Kansas Department of Wildlife and Parks, Pratt. vii + 106pp.
2007 Taggart, Travis W. A biological inventory of the Sunflower Electric Site near Holcomb, Kansas. Journal of Kansas Herpetology 23():11-16
2007 Taggart, Travis W., Joseph T. Collins, and Curtis J. Schmidt. Estimates of amphibian, reptile, and turtle mortality if Phostoxin is applied to 10,000 acres of prairie dog burrows in Logan County, Kansas. Kansas Department of Wildlife and Parks, Pratt. 5pp.
2008 Schmidt, Curtis J. Geographic distribution: Anaxyrus cognatus. Kansas. Journal of Kansas Herpetology (27):7
2010 Collins, Joseph T., Suzanne L. Collins, and Travis W. Taggart. Amphibians, Reptiles, and Turtles of Kansas Eagle Mountain Publishing., Provo, Utah. 400pp.
2010 Miller, Larry L. 2010 Investigation of the Checkered Garter Snake in Kansas with notes on other Amphibians, Reptiles, and Turtles encountered. Kansas Department of Wildlife and Parks, Pratt, Kansas. 31pp.
2011 Taggart, Travis W. Kansas Herpetological Society 2011 spring field trip to beheld in Chautauqua County. Journal of Kansas Herpetology (37):5-7
2011 Taggart, Travis W. Results of the Kansas Herpetological Society 2011 Summer Field Trip to Scott State Park Journal of Kansas Herpetology (39):2
2012 Rohweder, Megan R. Spatial conservation prioritization of Kansas for terrestrial vertebrates. Thesis. Fort Hays State University, Hays, Kansas. 151pp.
2013 Bass, Neil The Missouri River Fish and Wildlife Mitigation Project: For the river, for you, and for herps. Collinsorum 2(1/2):10-11
2013 Mardis, Dexter and Kevin Scott. 2013 Kansas Herpetofaunal Counts. Collinsorum 2(3/4):7
2013 Miller, Larry L. Wellington Lake Herpetological Survey. Collinsorum 2(1/2):12
2014 Taggart, Travis W. Results of the 2014 KHS Spring Field Trip to Barber County Collinsorum 3(2-4):11
2014 Taggart, Travis W. Results of the 2014 KHS summer field trip to Morton County and adjacent Colorado, New Mexico, and Oklahoma. Collinsorum 3(2-4):12
2015 Bass, Neil. Herpetological (Frog and Turtle) Inventories along the Missouri River in Kansas. Collinsorum 4(1):5-9
2015 Brown, Kasandra A. Occupancy Modeling Of Herpetofauna And Grassland Nesting Birds At Quivira National Wildlife Refuge. Thesis. Fort Hays State University, Hays, Kansas. 72pp.
2015 Taggart, Travis W. Spring Field Trip to the Greenhorn Limestone of Russell County. Collinsorum 4(3):2
2015 Taggart, Travis W. Summer Field Trip In The Harvey County Sandhills. Collinsorum 4(3):3
2016 Powell, Robert, Roger Conant, and Joseph T. Collins. Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America. Houghton Mifflin Harcourt, Boston. 494pp.
2016 Taggart, Travis W. Results of the KHS ‘Fall’ field trip to Barber County. Collinsorum 5(2-3):6-7
2016 Taggart, Travis W. Spring 2016 KHS field trip to Clark County was a soggy success. Collinsorum 5(2-3):2-3
2017 Snyder, Ariel Survey Of Anuran Chytrid (Batrachochytrium dendrobatidis) in Kansas and the Influence of Anuran Life History in Occurrence. Thesis. Fort Hays State University, Hays, Kansas. 53pp.
2017 Taggart, Travis W. and J. Daren Riedle. A Pocket Guide to Kansas Amphibians, Turtles and Lizards. Great Plains Nature Center, Wichita, Kansas. 69pp.
2020 Riedle, J. Daren. Revisiting Kansas Herpetological Society field trip and Herp Count data: Distributional patterns and trend data of Kansas amphibians and reptiles. Collinsorum 9(1):7-16
Account Last Updated:
9/18/2020 10:17:58 AM